Andrena (Micrandrena) dourada, Ratochwil, 2013

Ratochwil, 2013, Andrena (Micrandrena) dourada nov. sp. from Porto Santo, Madeira Archipelago, Portugal, Linzer biologische Beiträge 45 (1), pp. 755-774 : 762-764

2. Andrena (Micrandrena) dourada

F e m a l e. BL 6.89 mm (SD 0.24), WL 5.80 mm (SD 0.24); habitus Fig. 5.

C o l o u r. Head (Fig. 6): black, flagellum black (dark brown); mandible with distal half reddened. Mesosoma: black; femur tibia, basitarsus black; mediotarsi pars parte reddish brown; wings subhyalin, veins and pterostigma reddish brown, rarely brown; tegulae posterior part translucent, reddish brown, anterior part black. Metasoma: T1-4 black, with black to dark reddish brown depression zone, depression zone (T4) T5 translucent, reddish brown. In colour there is no difference from A. wollastoni .

P u b e s c e n c e. Head (Fig. 6): with yellowish-white hairs, in the paraocular area yellowish-white hairs existing from the malar area and overtop in contrast to A. wollastoni the line of the subantennal socket, near fovea only some few brownish hairs, hair length different; clypeus with shorter yellowish-white hairs, not dense, similar to supraclypeal area; distal end with two very long hairs; paraocular area with dense longer yellowish-white hairs (in general double length as clypeal hairs) near facial fovea with long brownish hairs; scapus and antennal socket with long brownish and yellowish hairs; genal area with dense long yellowish-white hairs similar to paraocular area; facial fovea with brown hairs; vertex with some yellowish-white longer hairs. Mesosoma (Fig. 7) only some brownish centrally located hairs; mesoscutum with lateral longer and denser yellowish/brownish hairs; scutellum with lateral shorter (half the length of mesoscutum) and denser yellowish/brownish hairs; mesepisternum with long yellowish-white hairs (double length of the hairs of the genal area); propodeal corbicula existent with long yellowish-white marginal hairs, some hairs in the centre; trochanteral and femoral flocculus well developed with yellowish-white hairs; tibial scopa with yellowish-white hairs; in contrast to A. wollastoni with fewer dorsoventral brown hair tips or basal fewer brownish hairs. Metasoma: tergites scarcely hairy, some longer hairs in the margin area. T4 with longer yellowish-white hairs, no distinct hair bands, but hair rows between tergite and tergite depression (T2-3 fragmentary developed); T5 laterally dense yellowish-white hairs, in the centre dense brownish hairs reaching to the pygidium.

S t r u c t u r e Head (Fig. 6): HL/HW = 0.83 (SD 0.24); HW:MsW:MtW = 1.1:1.1:1; vertex narrow, as wide as ocellar diameter, surface structure with granulate punctures; face above antennal fossae with longitudinal rugulae, interrugal space shiny; OOD:POD:OCD = 4.9:4.3:1; FL1:FL2:FL3 = 2.1:1:1; inner margin of eyes weakly converging; facial fovea FVL = 0.99 (SD 0.05), FVW = 0.19 (SD 0.02), FVL/FVW = 5.4 (SD 0.4); clypeus convex, shiny and smooth, without impunctate median line, shallow punctures (Pd = 28 μm), IS 14-84 μm, clypeus length: 0.56 mm (SD 0.04); labrum process (Fig. 8) in the rule liguliform (not emarginated rarely trapezoidliguliform), LBW = 0.08 mm (SD 0.02). Mesosoma (Fig. 7): mesonotum fine-grained, nearly unpunctured or weakly punctured; mesoscutum and scutellum smooth and shiny, very scattered punctures (Pd = 14 μm), mesoscutum with developed parapsidal lines; PSl = 0.86 mm (SD 0.03); propodeum basal very homogeneous only in some specimens rugose (Fig. 9), no lateral boundary line, PBAl = 0.37 mm (SD 0.01). Metasoma: smooth and shiny, very scattered punctures (Pd = 14 μm); posterior depression of (T1)T2-5.

Male. BL 5.93 mm (SD 0.23), WL 5.33 mm (SD 0.22).

C o l o u r.Nodifferences from female.

P u b e s c e n c e Head (Fig. 10): with dominantly white hairs, hair length slight different; clypeus with long white hairs, not dense, similar to supraclypeal and paraocular area (hairs yellowish-white); near facial fovea with long yellowish-white, only some brownish hairs; scapus and antennal socket with long yellowish-white hairs; genal area with dense long white hairs similar to paraocular area; vertex with yellowish-white hairs. Mesosoma: some yellowish long hairs; mesoscutum with lateral long and dense yellowish hairs; scutellum with shorter (half of length of mesoscutum) yellowish hairs; mesepisternum with long yellowish-white hairs (double length of the hairs of the genal area). Metasoma: tergites scarcely hairy, some longer hairs in the margin area. T4 with longer yellowish-white hairs, no distinct hair bands, but very fragmented hair rows between tergite and tergite depression (T2-3 fragmentary developed); T5 yellowishwhite hairs.

S t r u c t u r e. Head (Fig. 10): HL/HW = 0.83 (SD 0.05); HW:MsW:MtW = 1.1:1.1:1; vertex and face above antennal fossae similar to female; OOD:POD:OCD = 4.2:3.6:1; FL1:FL2:FL3 = 1.4:1:1; inner margin of eyes weakly converging; clypeus similar to female (Pd = 28 μm), IS 14-84(112) μm, clypeus length: 0.50 mm (SD 0.04); labrum process trapezoid, emarginated, left and right side thickened, LBW = 0.12 mm (SD 0.01). Mesosoma (Fig. 11): mesoscutum and scutellum similar to female (Pd = 14 μm); PSl = 0.76 mm (SD 0.03); propodeum structure similar to female, PBAl = 0.36 mm (SD 0.03). Metasoma: structure similar to female (Pd = 14 μm); posterior depression of (T1)T2-5. No differences in genital structure from A. wollastoni (Fig. 12).

D e r i v a t i o n o m i n i s A. dourada (substantival apposition). The Portuguese word "dourada" means "golden" or "aureate". Porto Santo is also named " Ilha dourada", "golden island", because of the white-yellow-brownish sand colours of the landscape. We can also build a bridge to the white-yellowish colour of the pubescense of this new species.

T y p e m a t e r i a l Holotype: PS12/117:, Porto Santo, near restaurant "Panorama" above Casinhas, 150 m, 33°04'8.47''N, - 16°19'14.57''E, yellow and white pan trap, 20.03.2012 ; Depository: Biology Centre of the Upper Austrian Provincial Museum Linz . Paratypes: (identity number, sex, locality, altitude above sea level, UTM coordinates, flower-visiting behaviour, date of collection): Females: PS12/47: 1, Porto Santo, Pedregal de Dentro , 220 m, 33°05'38.62''N, - 16°19'34.07''E, yellow pan trap, 20.03.2012 ; PS12/75: 1, Porto Santo, north of Serra de Dentro , 60 m, 33°05'15.36''N, - 16°18'33.15''E, yellow pan trap, 19.03.2012 ; PS12/110, PS12/111: 2, Porto Santo, Capela da Craça , above Vila Baleira, 160 m, 33°04'24.85''N, - 16°19'25.57''E, yellow pan trap, 20.03.2012 ; PS12/116: 1, Porto Santo, near restaurant "Panorama" above Casinhas, 150 m, 33°04'8.47''N, - 16°19'14.57''E, yellow and white pan trap, 20.03.2012 ; PS12/118, PS12/118: 2, Porto Santo, near Pico da Cabrita , 230 m, 33°05'42.18''N, - 16°19'04.49''E, white pan trap, 20.03.2012 ; PS12/137: 1, Porto Santo, north of Morenos , 90 m, 33°02'26.38''N, - 16°23'16.39''E, Leontodon taraxacoides (VILL.) MERAT , 20.03.2012 ; PS12/152: 1, Porto Santo, Campo de Baixo , near tennis court, 20 m, 33°02'50.07''N, - 16°21'23.35''E, Sinapis arvensis L., 20.03.2012 ; PS12/156: 1, Porto Santo, Capela de S. Pedro, 50 m, 33°02'44.85''N, - 16°21'43.82E, Rapistrum rugosum (L.) ALL., 20.03.2012. Males: PS12/43: 1, Porto Santo, southwest of Serra de Fora , 85 m, 33° 04'14.74"N, - 16°18'53.61"E, yellow pan trap, 18.03.2012 ; PS12/81: 1, Porto Santo, Vereda do Pico Branco , 310 m, 33°05'31.84''N, -16°18'17.16'', Calendula arvensis L., 19.02.2012 ; PS12/112, PS12/113: 2, Porto Santo, Capela da Craça , above Vila Baleira, 160 m, 33°04'24.85''N, - 16°19'25.57''E, yellow pan trap, 20.03.2012 .

Distribution and habitat characteristics

Individuals of A. dourada were detected on half of all investigated localities (10 from 21 localities, distributed all over the island); Fig. 13-16. So we can assume that A. dourada is widely distributed on Porto Santo. The range of altitudes reaches from sea level up to 310 m a.s.l. (highest point is Pico do Facho; 517 m a.s.l). Individuals of A. dourada were found primarily in the inframediterranean and lower thermomediterranean thermotypes; terminology according (CAPELO et al. 2004).

Flight time

There are not enough data available.

Flower visiting

So far, flower visits by females and males have been detected by A. Kratochwil and A. Schwabe on individuals of four plant taxa belonging to two plant families: Calendula arvensis L. ( Asteraceae ), Leontodon taraxacoides (VILL.) MERAT (Asteraceae) , Rapistrum rugosum (L.) ALL. ( Brassicaceae ), Sinapis arvensis L. ( Brassicaceae ). Additionally there is one observation made by COCKERELL (1922): Euphorbia spec. ( Euphorbiaceae ).

Differential diagnosis