Dziriblatta (Dziriblatta) bolivari Chopard, 1936a, 2019

Bohn, Horst, 2019, Revision of the genus Dziriblatta Chopard, 1936 (Blattodea, Ectobiidae, Ectobiinae) from North Africa, Spain, and the Macaronesian islands. I. The nine subgenera of the genus, Zootaxa 4610 (1), pp. 1-73 : 27

Dziriblatta (Dziriblatta) bolivari Chopard, 1936a , comb. nov.

Figs. 16A–I, 29.

Dziriblatta bolivari Chopard, 1936a: 159 .

Lobolampra bolivari: Chopard 1943: 35 , fig. 29.

Phyllodromica (Lobolampra) bolivari: Princis 1971: 1116 .

Dziriblatta opaca Chopard, 1936b: 159 ; Failla & Messina 1989: 34, figs. 3–8.

Diagnosis. In the male sex distinguished from all other Moroccan species of the subgenus by the relatively short and at least partly sclerotized anterior pouch of the T7 gland and the unique structure of the glandular pits.

Material studied.

Type material. Syntypes, 1♂, 1♀, Morocco, Tanger (terminalia of ♂, slide: Bo 1; of ♀, slide: Bo 2) [Ma 7]. ( MNHN) .

Additional material. Morocco. 1♀, Tanger , M. Escalera. ( ISNB) .— 1♂, 1♀, Rincon [Ma 8] ; 11♂, 7♀, 8L, Tanger , various dates and collectors ; 1♂, Targlitz , B.Seyel-Gomara (slide: Bo 199) [Ma 9] ; 1♀, Xauen (Ma 41) ; 1♀, Xauen , El Ajmas, Yebala, 9.–13.V.1932, M. Escalera ; 1♂, Yebel Dersa (Haus, Yebala) [Ma 1] V.1941, E. Morales ; 1♂, Yebel Lechab [= Jebel Lakraa], (El Ajmas, Yebala), V.1941 (slide: Bo 200) [Ma 10]. ( MNMS) .— 1♂, Tanger , “ Type ” ; 1♀, Maroc , Tanger ; 1L, Tanger, Olcese. ( MNHN) .— 1♂, ex L, Rif , btw. Ceuta & El-Biutz, 400 m, 29.III.1988, leg. B. & H.Bohn (slide: Ma 1a/2) ; 3♂, 1♀, 1L, Rif , 15 km W Bab-Berred, 1300 m, 1.IV.1988, leg. B. & H.Bohn (slides: ♂, Ma 5a/4; ♀, Ma 5a/10); Rif, Jbel Meggou (near Chefchaouèn), 1000 m, leg. B. & H.Bohn : 1♂, ex L, 6.IX.1984 (Ma 41); 2♂, 2♀, 1.IV.1988 (slides: ♂, Ma 41a/8; ♀, Ma 41a/8,9); 1♀, ex L, Rif , Jbel Tisouka (near Chefchaouèn), 1800 m, 6.IX.1984, leg. B. & H.Bohn (Ma 42) ; 1♀, Rif , Bge. Sedd-en-Nakhla, near Zinat (12 km S Tetouan), 450 m, 30.III.1988, leg. B. & H.Bohn (slide: Ma 43a/1) ; 1♀, Rif , Cap Spartel (near Tanger), 200 m, 29.III. / 7.IV.1988, leg. B. & H.Bohn (slide: Ma 47a/5); ex L : 1♂, Melilla Peninsula , Mt.Gourougou (S Melilla), 500 m, 15.IX.2000, leg. B. & H.Bohn (slide: Ma 70b/1) ; 1♂, 7♂, 19♀, Rif , E slope of Jbel Sougna (W Chefchaouèn), 750–900 m, 2./ 3.V.1997, leg. B. & H.Bohn (slides: ♂, Ma 123/1; ♀, Ma 123/2,3) ;? 1♀, Rif , Pays Beni Zerual, S slope of Lalla-Outka, 1100 m, 29.V.1997 (Ma 187) ;? 1♀, Rif , Pays Beni Zerual, Lalla-Outka, 1450-1550m, 29.V.1997 (Ma 188) ; 1♂, 1♀, Rif , 3.5 km SW Bab-Sidi-Kebat (8 km E Tahar-Souk), 1000 m, 11.IV.1998 (slide: ♂, Ma 222/1). (C. Bohn, ZSM). Spain . 3♂, 1♀, ex L: 1♂, 1♀, Prov. Granada , Sa. del Chaparral, Venta de Cabramontés (NW Otívar), 1000 m, 7.IV.1991, leg. B. & H.Bohn (slides: 2♂, Sp 209/1,2). (C. Bohn, ZSM)

Description. Size. Length of pronotum in male 2.08–2.40 (2.27 mean) mm, in female 2.46–2.82 (mean 2.66) mm.

Male structures. T7 with rather strongly produced latero-posterior corners (Figs. 16 A, B). Glandular pit of T7 similar to that of Dz. algerica , but smaller, occupying less than a third of the breadth of the tergite, excavations of the posterior wall occasionally deeper (Figs. 16A, I). The central pit is supplemented on each side by a large, in outline circular, moderately deep hole which is connected with the central pit by a relatively broad, shallow gutter. After filling the central pit the glandular secretions may find their way to the lateral pits via these gutters. However, the pit opening is not at the level of the surrounding tergite surface (as is the case in Dz. algerica ), instead, pit as a whole sunk to a considerably lower level. Thus, a larger secondary pit is formed, circumscribed anteriorly and laterally by the posterior edge of the pouch opening and the margins of the holes; posterior border less well demarcated, running along the line where the ascending surface posteriorly of the pit reaches the level of the proper tergite surface (arrow in Fig. 16B). Possibly, the secondary pit can be filled with secretions up to this level. Anterior lobes of the pouch much shorter than in Dz. algerica , scarcely longer than the length of the T7, conical, with a rather stout apex; wall usually well sclerotized. Glandular pores present on T2–5, usually in high numbers and relatively large size.

Colouration. In both sexes similar, variable, ground colour yellowish, thoracal nota to a various degree covered with differently sized light to dark brown markings (Figs. 16E–H), never as dark as in the male of the preceding species, abdominal tergites anteriorly with a dark transversal stripe, posteriorly often with dark dots and larger patches, abdominal sternites mainly dark.

Distribution. Northern Morocco between Cabo Espartel and Melilla peninsula (Fig. 29), on the latter found together with Dz. (Dz.) algerica at the same locality (Ma 70), and southern Spain (one locality). A report ( HARZ 1976) of the occurrence of Dz. algerica in southern Spain presumably refers to this species.