Bromeloecia magna Yau and Marshall, 2018

Yau, Tiffany & Marshall, Stephen A., 2018, A revision of the genus Bromeloecia Spuler (Diptera: Sphaeroceridae: Limosininae), Zootaxa 4445 (1), pp. 1-115 : 22-26

Bromeloecia magna Yau and Marshall sp. n.

(Figs. 3.1–3.14)

Description: Body length 1.2–1.7 mm (males), 1.5–2.1 mm (females). Generally similar to B. abundantia , but the colour contrast between the yellowish gena and the dark brown thorax is more pronounced.

Wing: Distance between radial fork and r-m 6x length of r-m; dm-cu 2x length of r-m and distance between crossveins 2.3x length of dm-cu.

Male abdomen and terminalia: S5 with a pair of posteromedial lobes, lobes posteroventrally with medially directed spicules; a row of strong setae anterior to lobes; row longer than width of modified posteromedial part of sclerite. Cercus laterally compressed, uniform in width, apex distinctly bent anteriorly (almost 90°). Surstylus ventrolaterally with thumb-like process and ventrally with smaller tooth-like protuberance. Distiphallus LFS sheetlike, apically long and tapered; FDS form a posteriorly bent tongue; SDS weakly sclerotized, spatulate.

Type specimens: Holotype: ♂: COSTA RICA: San José: Zurquí de Moravia , 10°02ʹ58ʺN, 84°00ʹ57ʺW, 1600 m, Malaise trap, 9–19 Oct 2012, INB0004357594 ( INBC) .

Paratypes: COSTA RICA : Cartago: Tapantí Natl. Pk., Ranger Stn. 09°45ʹ22ʺN, 083°47ʹ05ʺW, 1200 m, human dung, hand & traps, 9–12 Oct 1999, Buck & Marshall (15♂, 7♀, INBC, DEBU) ; Tapantí Natl. Pk., 09°42ʹ18ʺN, 083°43ʹ04ʺW, 1550 m, pans in fallen tree, 7–12 Oct 1999, Marshall & Buck (1♂, DEBU); Tapantí Natl. Pk. , 09°42ʹ18ʺN, 083°43ʹ04ʺW, 1650–1750 m, sweeping trail, 7 Oct 1999, M. Buck (1♂, DEBU) ; Tapantí Natl. Pk., Arboles Caidos Trail , on foliage, 1200 m, 11 Oct 1999, Marshall & Buck (1♀, DEBU) ; Guanacaste: Cacao Station , 1250 m, dung trap, 12–15 Feb 1996, S. A. Marshall (1♂, DEBU) ; Heredia: La Virgen , 16 km SSE, 10°16ʹN, 084°05ʹW, 1050–1150 m, transect, 9–21 Apr 2001 (1♂, DEBU) ; La Virgen , 16 km SSE, 10°16ʹN, 084°05ʹW, 1050–- 1150 m, 9 Mar 2001 (1♀, DEBU) ; ditto, 21 May 2001 (1♀, DEBU) ; Puntarenas: Monteverde Biol. Res. , 10°19ʹ07ʺN, 084°48ʹ03ʺW, 1500 m, tree fall sweep & pans, 14 Jun 2000, M. Buck (1♂, DEBU) ; Monteverde Biol. Res. , 10°19ʹ07ʺN, 084°48ʹ03ʺW, cloud forest, 1500 m, 11–13 Jun 2000, S. A. Marshall (5♂, 3♀, INBC, DEBU) ; Monteverde Biol. Res. , 10°19ʹ07ʺN, 084°48ʹ03ʺW, 1500 m, 13 Jun 2000, S. A. Marshall (1♂, DEBU) ; Monteverde Biol. Stn. , 1500 m, dung RET, 26 May 1998, S. A. Marshall (2♀, DEBU) ; Monteverde , cloud forest, 1500 m, 29 Feb 1980, W. R. Mason (1♀, DEBU) ; Monteverde , cloud forest, 1500 m, dung trap, 19–25 Aug 1993, E. R. Barr (1♀, DEBU) ; Monteverde , dung RET nr. station, 26 May 1998, S.A. Marshall (1♀, DEBU) ; Monteverde , 1520 m, FIT, 23–30 Jul 1983, D. H. Lindeman (1♀, DEBU) ; Monteverde , rainforest, 1500 m, 29 Feb 1980, Mason & Wood (1♀, DEBU) ; San José: Zurquí de Moravia , tower path, 10°02ʹ58ʺN, 084°00ʹ57ʺW, 1600 m, Malaise trap, 30 Aug–6 Sep 2013 (1♂, 1♀, INBC) ; ditto, 3–10 Aug 2013 (1♀, INBC) ; Zurquí de Moravia, North pasture, 10°02ʹ58ʺN, 084°00ʹ57ʺW, 1600 m, Malaise trap, 3–11 Jan 2013 (1♀, INBC) . PANAMA : Chiriquí: La Fortuna Dam , 1000 m, 5–6 Jul 1981, B. Gill (1♀, DEBU)

Etymology: This species is the largest, most heavy-bodied species of Bromeloecia . Thus, the name refers to the body size (Latin ‘magnus’: large, great).

Comments: B. magna is found at relatively high elevations in the mountain ranges of Costa Rica and northern Panama. It is the largest species of Bromeloecia . Although B. magna and B. abundantia differ widely in body size, they are very similar in external characters (to differentiate the two, see comments below the description of B. abundantia ). Male cerci of B. magna are always flexed inwards in dry specimens.