Ammophila kowalczyki Olszewski, 2020

Olszewski, Piotr & Danilov, Yuriy N., 2020, New species and records of the genus Ammophila W. Kirby, 1798 (Hymenoptera Sphecidae), Zootaxa 4878 (1), pp. 189-194 : 190

Ammophila kowalczyki Olszewski sp. nov.

Fig. 1 A–M

Etymology. The species is named after my friend Jan Krzysztof Kowalczyk from Gdynia, Poland, an expert in Polish Aculeata.

Type material. Holotype, ♂, IRAN: Kerman, Gohargaz [N30.43753, E57.21039], 2020 m a.s.l., 9 May 2017, leg. Iran Expedition IV, in coll. NHMUK ; Paratype, ♀, Iran: Kerman, Gohargaz [N30.43753, E57.21039], 2020 m a.s.l., 13 May 2017, leg. Iran Expedition IV, in coll. NHMUK .

Diagnosis. In Ammophila kowalczyki , the metasomal apex is black with a metallic lustre, and the propodeal enclosure is wholly glabrous and transversely striated ( Fig. 1I ). The pronotal collar and scutum are distinctly ridged transversely ( Fig. 1G, H ). The male shares a blunt tooth on the hypostomal carina with A. abnormis Dollfuss, 2013 , A. horni von Schulthess, 1927 and A. punctata F. Smith, 1856 ; of these species, only A. abnormis has a wholly glabrous propodeal enclosure, and it shares a similarly shaped clypeus with the new species.

The male of A. kowalczyki differs by the gonostyle with distinct, long preapical spines, the shape of the cuspis (wide and not visibly tapering) and the shape of the penis valve ( Fig. 1E ).

In A. abnormis , the gonostyle has at most a few short preapical spines and a slender, characteristically tapering cuspis. The penis valve has no preapical spines.

Both sexes of A. kowalczyki resemble A. gussakovskii Dollfuss, 2013 in having a distinct patch of appressed silvery setae on the mesopleuron and a glabrous propodeal enclosure. Also, the pronotal collar and scutum are transversely striate and the gastral apex has a metallic lustre. The male of A. kowalczyki differs in having the hypostomal carina with a tooth near the mandibular socket. The female of A. kowalczyki differs by the appressed silvery setae over the whole mesopleuron except for its upper part.

Description. Female. Body length 16 mm. Head ( Fig. 1C ). Clypeal disk sparsely punctured medially. Clypeus weakly elongate ventrally, with appressed silvery setae in basal part. Median lobe of clypeus emarginate ( Fig 1C ). Subantennal sclerite with appressed silvery setae. Gena, vertex and frons with long separate white setae and mandible with long separate dark setae. Supra-antennal lamellate projection absent. Frons and vertex dull. F1: F2 = 6.2: 3.2.

Mesosoma. Pronotum not elongate. Propodeal enclosure glabrous, obliquely striated. Mesopleuron with appressed silvery setae concealing sculpture (except for its upper part), metapleuron irregularly rugose, striated. Scutellum and metanotum longitudinally striated. Propleuron with dense appressed silvery setae and long separate white setae.

Legs. Coxae with dense appressed silvery setae dorsally. Claws without basal tooth.

Fore wing with three submarginal cells. Submarginal cell III not petiolate. Marginal cell with rounded apex.

Colouration. Head, mesosoma, legs entirely black. Wings hyaline; venation dark brown. T4 and 2/5 of T5 laterally, tegula, petiole, T1–T3, and S1–S5 red. Other parts of metasoma black with metallic lustre.

Male. Head, mesosoma, and metasoma as in female except for the following: Body length 16 mm. Head ( Fig. 1D ). Clypeus elongate, medially with one prominent teeth ( Fig. 1D ), elevated in lateral view, with dense appressed silvery setae concealing sculpture except for apical margin. Hypostomal carina with tooth near mandibular socket ( Fig. 1J ). Mandible with sparse long white and dark setae. F1: F2 = 5.5: 3.2. Scutum finely transversely striated anteriorly, distinctly obliquely striated posteriorly (in male more so than in female). Male genitalia (as illustrated ( Fig. 1 E–F). Gonostyle with long preapical spines ( Fig. 1E ), penis valve without preapical spines ( Fig. 1E ). Cuspis wide, not visibly tapering ( Fig. 1F ). Colouration. T4, ⅔ of T5, ½ of T6, ¼ of T7 laterally, pronotal lobe, petiole, T1–T3, and S1–S7 red. Other parts of metasoma black with metallic lustre.

Figures

FIGURE 1. Ammophila kowalczyki sp. nov. Holotype ♂. Paratype ♀: (A) ♀, habitus, lateral view. (B) ♂, habitus, lateral view. (C) ♀, head, frontal view. (D) ♂, head, frontal view. (E) ♂, genitalia, lateral view. (F) ♂, genitalia, oblique ventral view. (G) ♀, pronotum, scutum, dorsal view. (H) ♂, pronotum, scutum, dorsal view. (I) ♀, propodeal enclosure, dorsal view. (J) ♂, hypos- tomal carina, lateral view. (K) ♂, metasoma apex, lateral view. (L) ♀, metasoma apex, lateral view. (M) ♀, fore and hind wings, dorsal view.