Everettia dominiki, Liew & Schilthuizen & Vermeulen, 2009

Liew, Thor-Seng, Schilthuizen, Menno & Vermeulen, Jaap Jan, 2009, Systematic revision of the genus Everettia Godwin-Austen, 1891 (Mollusca: Gastropoda: Dyakiidae) in Sabah, northern Borneo, Zoological Journal of the Linnean Society 157 (3), pp. 515-550 : 542-543

publication ID	https://doi.org/ 10.1111/j.1096-3642.2009.00526.x
DOI	https://doi.org/10.5281/zenodo.10545936
persistent identifier	https://treatment.plazi.org/id/03F9878A-FF94-E81F-0D02-89CB22E1FA3E
treatment provided by	Felipe
scientific name	Everettia dominiki
status	sp. nov.

Treatment

EVERETTIA DOMINIKI View in CoL SP. NOV.

Types: MALAYSIA: State of Sabah: Kota Marudu District. North-eastern part of Kinabalu Park. At 1600 m alt. along the trail from Serinsim Substation to Mount Nombuyukon No (6°15 ′ N, 116°40 ′ E). Collected by T.- S. Liew. Date: 9.xi.2005, holotype, BOR/ MOL 5360, Figure 12C View Figure 12 ; six paratypes, three in BMNH 20080635 and three in SP 13067 GoogleMaps .

Etymology: This species is named after Dominik Tinggoh, staff member at Kinabalu Park, who dedicatedly assisted the first author during the intensive malacological survey on Mount Kinabalu in 2005.

Material examined: SABAH – Mount Kinabalu: BOR/ MOL 1314, southern slope, 1450 m alt., vi.2002; BOR/ MOL 2691, southern slope, 2865 m alt., x.2003; BOR/MOL 2697, southern slope, 2267 m alt., x.2003; BOR/MOL 2698, southern slope, 2460 m alt., x.2003; BOR/MOL 5380, southern slope, 2112 m alt., iv.2005; BOR/MOL 5381, southern slope, 2404 m alt., iv.2005; BOR/MOL 5383, southern slope, 2096 m alt., ix.2005; BOR/MOL 5385, southern slope, 2628 m alt., iv.2005; BOR/MOL 5386, southern slope, 2288 m alt., iv.2005; BOR/MOL 5388, southern slope, 2308 m alt., xii.2004; BOR/MOL 5390, southern slope, 2526 m alt., xii.2004; BOR/MOL 5392, southern slope, 2552 m alt., ix.2005; BOR/MOL 5396, southern slope, iv.2006; BOR/MOL 5399, southern slope, 3154 m alt., i.2005; BOR/MOL 5400, southern slope, 2896 m alt., ix.2005; BOR/MOL 5401, southern slope, 3024 m alt., ix.2005; BOR/MOL 5402, southern slope, 2896 m alt., xii.2004; BOR/MOL 5403, southern slope, 3088 m alt., ix.2005; BOR/MOL 5405, southern slope, 2528 m alt., iv.2005; BOR/ MOL 5408, southern slope, 1948 m alt., iv.2005; BOR/ MOL 5410, southern slope, 2092 m alt., iv.2005; BOR/MOL 5412, southern slope, 2040 m alt., iv.2005; BOR/MOL 5413, southern slope, 2120 m alt., iv.2005; BOR/MOL 5414, southern slope, 2120 m alt., iv.2005; BOR/MOL 5417, southern slope, 1784 m alt., ix.2005; BOR/MOL 5419, southern slope, 2484 m alt., xii.2004; BOR/MOL 5398, western slope, 2640 m alt., ix.2005; BOR/MOL 5409, western slope, 1968 m alt., ix.2005; BOR/MOL 5422, western slope, 2576 m alt., ix.2005; BOR/MOL 5387, eastern slope, 2812 m alt., iv.2005; BOR/MOL 5389, eastern slope, 2676 m alt., iv.2005; BOR/MOL 5391, eastern slope, 2132 m alt., iv.2005; BOR/MOL 5404, eastern slope, 1992 m alt., iv.2005; BOR/MOL 5411, eastern slope, 1952 m alt., iv.2005; BOR/MOL 5387, eastern slope, 2812 m alt., iv.2005; BOR/MOL 5415, eastern slope, 2300 m alt., iv.2005; BOR/ MOL 5416, eastern slope, 3112 m alt., iv.2005; BOR/ MOL 5418, eastern slope, 2992 m alt., iv.2005; BOR/MOL 5420, eastern slope, 3064 m alt., iv.2005; BOR/MOL 5421, eastern slope, 2188 m alt., iv.2005; BMNH 20080208, southern slope, Mesilau, vi.1967; BMNH 20080209, southern slope, Mesilau, vi.1967. Mount Tambuyukon: BOR/MOL 5383, eastern slope, 2288 m alt., ix.2005; BOR/MOL 5384, eastern slope, 2080 m alt., ix.2005; BOR/MOL 5395, eastern slope, ix.2005; BOR/MOL 5397, eastern slope, 1584 m alt., ix.2005; BOR/MOL 5406, eastern slope, 1952 m alt., ix.2005; BOR/MOL 5407, eastern slope, 3260 m alt., ix.2005.

Diagnostic characteristics: Shell surface with dense and regularly radial ribs.

Description: Shell ( Figs 3F View Figure 3 , 12C View Figure 12 ): medium-sized, rather thin, brownish, spire moderately to highly elevated, outer whorls shouldered below the suture. Periphery rounded. Above the periphery, shell silky with fine, dense, and regular radial corrugation, c. 20 wrinkles per mm. Below the periphery, shell has corrugation that extends from above the periphery and dense shallow spiral grooves, but some individuals with only faded spiral grooves on the bottom surface of the shell or entirely without these. Height up to 8.3 mm; width up to 10.1 mm; diameter of the first three whorls 0.7, 0.9, and 1.2 mm, respectively; number of whorls up to four and a half; height aperture up to 4.7 mm; width aperture up to 5.5 mm. Genitalia ( Fig. 13B View Figure 13 ): maximum length from genital opening to the end of dart-sac (before the visible gland tubules) up to 7 mm. The penis and dart-sac have thin muscular walls but vagina has a very thin transparent wall. P, V, GO, BC, and DS arranged around the short genital atrium, with limited space between the opening of each of the genital structures. BC long, almost as long as DS. Animal ( Fig. 8C View Figure 8 ): black bands on either side of the tentacles, which are separated by a white band that occupies the space between the eye tentacles and extends backward to the end of the mantle and downward to the foot fringe. However, in a few rare cases the animal’s head is uniformly black. Other parts of the animal are almost entirely black. Some blackish smears and bright markings are irregularly arranged on the brownish mantle. The mantle in the last half whorl is covered by sparse, irregular black and bright markings.

Distribution and habitat: Montane primary forest 1500–3300 m (common). Sabah: Mount Kinabalu (endemic?) ( Fig. 9C View Figure 9 ).

Remarks: The two morphs of this species – with and without spiral grooves on the underside of the shell, occur sympatrically but the one with these spiral grooves can mainly be found between 2000–2500 m. However, genetic data do not support the monophyly of these two morphs.