Everettia monticola, Liew & Schilthuizen & Vermeulen, 2009

Liew, Thor-Seng, Schilthuizen, Menno & Vermeulen, Jaap Jan, 2009, Systematic revision of the genus Everettia Godwin-Austen, 1891 (Mollusca: Gastropoda: Dyakiidae) in Sabah, northern Borneo, Zoological Journal of the Linnean Society 157 (3), pp. 515-550 : 541-542

publication ID	https://doi.org/ 10.1111/j.1096-3642.2009.00526.x
DOI	https://doi.org/10.5281/zenodo.5492205
persistent identifier	https://treatment.plazi.org/id/03F9878A-FF95-E818-0FB4-8D2121F7FA53
treatment provided by	Felipe
scientific name	Everettia monticola
status	sp. nov.

Treatment

EVERETTIA MONTICOLA View in CoL SP. NOV.

Types: MALAYSIA: State of Sabah: Kota Belud District. Southern part of Kinabalu Park. Near ‘ Villosa Shelter’ at 2900 m alt. along the Summit trail of Mount Kinabalu (6°3 ′ N, 116°33 ′ E). Collected by Tachani. Date: x.2003, holotype, BOR/MOL 2691, Figure 12B View Figure 12 ; three paratypes, two in BMNH 20080634 and one in SP 13066 GoogleMaps .

Etymology: This species is named monticola for its montane distribution.

Material examined: SABAH – Mount Trusmadi: BOR/ MOL 1310, 1600 m alt., iii.2002; BOR/MOL 1138, 880 m alt., x.2001. Crocker Range: BOR/MOL 1312, Ulu kimanis, 1200 m alt., vi.2003; BOR/MOL 1313, Mahua, 1000 m alt., iii.2002; BOR/MOL 1131, Mahua, 1000 m alt., v.2001; BOR/MOL 1136, Mahua, 1000 m alt., vii.2001. Kinabatangan: BOR/MOL 1411, Batu Tomanggong kecil, iv.2002. Meliau Range: BOR/MOL 3214, 640 m alt., viii.2004; BOR/MOL 3215, 650 m alt., viii.2004. Mount Kinabalu: BOR/MOL 5361, southern slope, 1552 m alt., ix.2005; BOR/MOL 5364, southern slope, iii.2005; BOR/MOL 5368, southern slope, 2156 m alt., iv.2005; BOR/MOL 5369, southern slope, 1784 m alt., ix.2005; BOR/MOL 5374, southern slope, 1700 m alt., ii.2005; BOR/MOL 5376, southern slope, iii.2005; BOR/MOL 5378, southern slope, 2112 m alt., iv.2005; BOR/MOL 5365, western slope, 1776 m alt., xi.2005; BOR/MOL 5366, western slope, 1152 m alt., xi.2005; BOR/MOL 5370, western slope, 2480 m alt., xi.2005; BOR/MOL 5371, western slope, 1152 m alt., xi.2005; BOR/MOL 5377, western slope, 1496 m alt., xi.2005; BMNH 20080205, southern slope, Mesilau, vi.1967; BMNH 20080206, southern slope, Mesilau, vi.1967; BMNH 20080207, southern slope, Mesilau, vi.1967. Mount Tambuyukon: BOR/MOL 5372, eastern slope, 1144 m alt., ix.2005; BOR/MOL 5373, eastern slope, 1880 m alt., ix.2005; BOR/MOL 5375, eastern slope, 1584 m alt., ix.2005. Mount Nombuyukon: BOR/ MOL 5360, north-eastern slope, 1204 m alt., xi.2005; BOR/MOL 5362, north-eastern slope, 1070 m alt., xi.2005; BOR/MOL 5363, north-eastern slope, 1680 m alt., xi.2005; BOR/MOL 5367, north-eastern slope, 1680 m alt., xi.2005; BOR/MOL 5379, north-eastern slope, xi.2005.

Diagnostic characteristics: This species is without distinct characteristics on the shell, except for the moderately elevated spire, in which it differs from other Everettia species of similar size.

Description: Shell ( Figs 3G View Figure 3 , 12B View Figure 12 ): medium-sized, rather thin, brownish, spire moderately elevated, outer whorls shouldered below suture. Periphery rounded. Above the periphery, smooth but sometimes with irregularly placed oblique wrinkling that is slightly stronger towards the suture. Below the periphery, shell is with or without fine, shallow spiral grooves. Height up to 7.0 mm; width up to 11.9 mm; diameter of the first three whorls 0.7, 0.5, and 1.0 mm, respectively; number of whorls up to five; height aperture up to 5.4 mm; width aperture up to 6.5 mm. Genitalia ( Fig. 13B View Figure 13 ): maximum length from genital opening to the end of dart-sac (before the visible gland tubules) up to 7 mm. The penis and dart-sac have thin muscular walls, and vagina has very thin transparent wall. P, V, GO, BC, and DS arranged around the short genital atrium, with limited space between the openings of each of the genital structures. BC long, almost same length as DS. Animal ( Fig. 8C View Figure 8 ): black bands on either side of the tentacles, which are separated by a white band that occupies the space between the eye tentacles and extends backward to the end of the mantle and downward to the foot fringe, although in some animals, the head around the tentacles is uniformly black but a white band appears near the end of mantle. Other parts of the animal are covered by regularly spaced oblique dotted belts or a grey smear. The mantle is brownish or greyish and in the last half whorl with sparse irregular black and bright markings.

Distribution and habitat: Tropical lowland and montane primary forest 700–2500 m (common between 1000–1500 m). Sabah: Crocker Range, Trusmadi Range, Mount Kinabalu ( Fig. 9A View Figure 9 ).

Remarks: The shell of this species is similar to E. robusta , which was described from Borneo ( Gude, 1917). However, it is impossible to determine where in Sabah (if at all) the type was collected.