Everettia safriei, Liew & Schilthuizen & Vermeulen, 2009

Liew, Thor-Seng, Schilthuizen, Menno & Vermeulen, Jaap Jan, 2009, Systematic revision of the genus Everettia Godwin-Austen, 1891 (Mollusca: Gastropoda: Dyakiidae) in Sabah, northern Borneo, Zoological Journal of the Linnean Society 157 (3), pp. 515-550 : 540

publication ID	https://doi.org/ 10.1111/j.1096-3642.2009.00526.x
DOI	https://doi.org/10.5281/zenodo.5492199
persistent identifier	https://treatment.plazi.org/id/03F9878A-FF96-E81A-0D08-8DD827E4FC4A
treatment provided by	Felipe
scientific name	Everettia safriei
status	sp. nov.

Treatment

EVERETTIA SAFRIEI View in CoL SP. NOV.

Types: MALAYSIA: State of Sabah: Ranau District. Southern part of Kinabalu Park. At 2300 m alt. along Kotal’s route from Bukit Babi to the eastern ridge of Mount Kinabalu (6°3 ′ N, 116°36 ′ E). Collected by T.- S. Liew, J. Lapidin, Safrie, and Jasilin. 24.iv.2005, holotype, BOR/MOL 5307, Figure 12D View Figure 12 ; six paratypes, three in BMNH 20080631 and three in SP 13063 GoogleMaps .

Etymology: This species is named after Safrie Hatimin, staff member at Kinabalu Park, who dedicatedly assisted the first author during the intensive malacological survey on Mount Kinabalu in 2005.

Material examined: SABAH – Mount Kinabalu: BOR/ MOL 2671, southern slope, 3330 m alt., x.2003; BOR/ MOL 2672, southern slope, 3330 m alt., x.2003; BOR/MOL 2682, southern slope, 3080 m alt., x.2003; BOR/MOL 2683, southern slope, 3080 m alt., x.2003; BOR/MOL 2687, southern slope, 3080 m alt., x.2003; BOR/MOL 5302, southern slope, 3024 m alt., ix.2005; BOR/MOL 5306, southern slope, 3089 m alt., xii.2004; BOR/MOL 5303, eastern slope, 3264 m alt., vi.2005; BOR/MOL 5304, eastern slope, 3352 m alt., vi.2005; BOR/MOL 5305, eastern slope, 3432 m alt., vi.2005; BOR/MOL 5308, eastern slope, 3416 m alt., vi.2005; BOR/MOL 5309, eastern slope, 2924 m alt., iv.2005; BOR/MOL 5310, eastern slope, 3336 m alt., vi.2005.

Diagnostic characteristics: The numerous, irregular, very fine, and discontinuous spiral threads on the upper surface of the shell differentiate this species from individuals of Everettia dominiki with similar shell shape and size.

Description: Shell ( Figs 10C View Figure 10 , 12D View Figure 12 ): small, rather thin, greenish brown, moderately elevated, outer whorls rounded. Periphery rounded; spire almost flat. Above the periphery, protoconch smooth (sometimes with inconspicuous, densely placed spiral threads); shell with distinct oblique wrinkling; there are very fine, irregular and discontinuous spiral threads above the wrinkling. Below the periphery, shell is almost smooth. Height up to 4.8 mm; width up to 8.3 mm; diameter of the first three whorls 0.75–0.8, 0.55–0.60, and 0.95–1.00 mm, respectively; number of whorls up to five and one-quarter; height aperture up to 3.6 mm; width aperture up to 4.5 mm. Genitalia ( Fig. 13C View Figure 13 ): maximum length from genital opening to the end of dart-sac (before the visible gland tubules) up to 4.5 mm. The penis, dart-sac, and vagina with very thin transparent walls. P, V, GO, BC, and DS arranged around the short genital atrium, with limited space between the openings of each of the genital structures. BC short, about half of total DS length. Animal ( Fig. 8D View Figure 8 ): the head of animal head is uniformly black. The mantle is brownish and in the last half whorl has sparse irregular black and bright markings.

Distribution and habitat: Upper montane primary and coniferous forest, 2700–3500 m alt. Sabah: Mount Kinabalu (eastern and southern ridges only), endemic ( Fig. 9A View Figure 9 ).

Remarks: This species occurs sympatrically with E. safriei , to which it is also phylogenetically close. However, both species maintain their distinct and unique shell characteristics.