Everettia layanglayang, Liew & Schilthuizen & Vermeulen, 2009

Liew, Thor-Seng, Schilthuizen, Menno & Vermeulen, Jaap Jan, 2009, Systematic revision of the genus Everettia Godwin-Austen, 1891 (Mollusca: Gastropoda: Dyakiidae) in Sabah, northern Borneo, Zoological Journal of the Linnean Society 157 (3), pp. 515-550 : 534-535

publication ID	https://doi.org/ 10.1111/j.1096-3642.2009.00526.x
DOI	https://doi.org/10.5281/zenodo.5492189
persistent identifier	https://treatment.plazi.org/id/03F9878A-FF9C-E816-0FB7-89F12171FF30
treatment provided by	Felipe
scientific name	Everettia layanglayang
status	sp. nov.

Treatment

EVERETTIA LAYANGLAYANG View in CoL SP. NOV.

Types: MALAYSIA: State of Sabah: Ranau District. Southern part of Kinabalu Park. At 3112 m alt. along Kotal’s route from Bukit Babi to the eastern ridge of Mount Kinabalu (6°3 ′ N, 116°36 ′ E). Collected by T.- S. Liew, J. Lapidin, Safrie, and Jasilin. 24.iv.2005; holotype, BOR/MOL 4439, Figure 6F View Figure 6 GoogleMaps ; MALAYSIA: State of Sabah: Ranau District. Southern part of Kinabalu Park. At 2200 m alt. along Kotal’s route from Bukit Babi to the eastern ridge of Mount Kinabalu (6°3 ′ N, 116°36 ′ E). Collected by T.- S. Liew, J. Lapidin, Safrie, and Jasilin. 24.iv.2005; two paratypes, one in BMNH 20080626 and one in SP 13058 GoogleMaps .

Etymology: This species is named layanglayang after the location on Mount Kinabalu where living snails were first found with their unique mantle pigmentation that differs from other Everettia species.

Material examined: SABAH – Mount Kinabalu: BOR/ MOL 941, southern slope, 3200 m alt., xi.2001; BOR/ MOL 950, southern slope, 1400 m alt., vii.2001; BOR/MOL 920, southern slope, 1400 m alt., iii.2000; BOR/MOL 971, southern slope, 1400 m alt., i.2001; BOR/MOL 1379, southern slope, 1400 m alt., vi.2002; BOR/MOL 2699, southern slope, 2460 m alt., xi.2003; BOR/MOL 4246, southern slope, 1500 m alt., iv.2006; BOR/MOL 4429, southern slope, 2300 m alt., xii.2004; BOR/MOL 4430, southern slope, 2308 m alt., xii.2004; BOR/MOL 4431, eastern slope, iv.2005; BOR/MOL 4432, southern slope, 1600 m alt., iii.2005; BOR/MOL 4433,eastern slope, 2300 m alt., iv.2005; BOR/MOL 4435, southern slope, 3221 m alt., i.2005; BOR/MOL 4436, southern slope, 2092 m alt., iv.2005; BOR/MOL 4437, southern slope, 2120 m alt., iv.2005; BOR/MOL 4438, eastern slope, 2412 m alt., iv.2005; BOR/MOL 4441, southern slope, 2188 m alt., iv.2005; BOR/MOL 4442, eastern slope, 2292 m alt., iv.2005; BOR/ MOL 4443, southern slope, 2288 m alt., iv.2005; BOR/ MOL 4444, southern slope, 2000 m alt., iv.2005; BOR/MOL 4445, eastern slope, iv.2005; BOR/MOL 4446, western slope, 1776 m alt., xi.2005; BOR/MOL 4447, eastern slope, 2060 m alt., iv.2005; BOR/ MOL 4448, southern slope, 1784 m alt., ix.2005; BOR/ MOL 4449, southern slope, 2212 m alt., iv.2005; BOR/MOL 4450, southern slope, 2096 m alt., ix.2005; BOR/MOL 4451, southern slope, 1616 m alt., iii.2005; BOR/MOL 4452, southern slope, 1700 m alt., ix.2005; BOR/MOL 4453, eastern slope, 1992 m alt., ix.2005; BOR/MOL 4454, southern slope, 1720 m alt., ix.2005; BOR/MOL 4455, southern slope, 2108 m alt., ii.2005; BOR/MOL 4456, eastern slope, 1952 m alt., iv.2005; BOR/MOL 4457, southern slope, iii.2005; BOR/MOL 4458, eastern slope, 2280 m alt., iv.2005; BOR/MOL 4459, southern slope, 2040 m alt., iv.2005; BOR/MOL 4460, southern slope, 2244 m alt., iv.2005; BOR/MOL 4461, eastern slope, 2604 m alt., iv.2005; BOR/MOL 4462, southern slope, 2244 m alt., iv.2005; BOR/MOL 4463, southern slope, 2528 m alt., iv.2005; BOR/MOL 4464, southern slope, 2078 m alt., ii.2005; BOR/MOL 4460, western slope, 1776 m alt., xi.2005; BOR/MOL 4466, southern slope, 1700 m alt., ii.2005; BOR/MOL 4467, southern slope, 1784 m alt., ix.2005; BOR/ MOL 4468, southern slope, 2560 m alt., iv.2005; BOR/ MOL 4469, southern slope, 2244 m alt., iv.2005; BOR/MOL 4470, southern slope, 1723 m alt., xii.2004; BOR/MOL 4471, southern slope, 2356 m alt., iv.2005; BOR/MOL 4472, southern slope, 2628 m alt., iv.2005; BOR/MOL 4473, eastern slope, 2412 m alt., iv.2005; JJ 1190, southern slope, 1500 m alt., viii, 1986; JJ 13022, southern slope, 2100 m alt., vii.2005; SP 12504, southern slope, 1800 m alt., iv.2005; SP 12924, southern slope, Marai-parai, v.2007; BMNH 20080204, southern slope, 1900 m alt., viii.1967. Crocker Range: BOR/MOL 968, Mount Emas, vii.2000; SP 12907, Mount Alab, 1800 m alt., iii.2007. Mount Trusmadi: BOR/MOL 1384, 1600 m alt., iii.2002; BOR/MOL 1386, 1600 m alt., iii.2002.

Diagnostic characteristics: Uniform pinkish, reddish brown or black animal with unique and regular spot markings on the mantle.

Description: Shell ( Figs 6F View Figure 6 , 3C, E View Figure 3 ): large, rather thin, yellowish brown to reddish brown. Spire moderately elevated, outer whorls almost rounded below the suture. Periphery almost rounded. Above the periphery, shell silky with very weak oblique wrinkling, slightly stronger towards the suture. Amongst this wrinkling, very fine granulation (40¥ magn.), is arranged obliquely. Below the periphery, shell has fine, densely placed spiral grooves. Height up to 10.7 mm; width up to 17.7 mm; diameter of the first three whorls 0.9–1.1, 0.8–1.0, and 1.4–1.6 mm, respectively; number of whorls up to five and one-quarter; height aperture up to 7.4 mm; width aperture up to 9.3 mm. Genitalia ( Fig. 7F View Figure 7 ): maximum length from genital opening to the end of dart-sac (before the visible gland tubules) up to 14 mm. The penis, dart-sac, and vagina with thick muscular walls. P, V arranged near the GO then followed by BC and DS where there is little space in the atrium between the openings of P + V and BC + DS. BC almost same as DS length. Animal ( Figs 4E View Figure 4 , 8B View Figure 8 ): the whole animal head is uniformly pinkish, reddish brown, or black. The mantle is darker grey and covered by sparsely distributed tiny brighter spots. These brighter spots become bigger and regularly arranged at the last half whorl.

Distribution and habitat: Primary forests. Sabah: Mount Kinabalu (1800–2600 m) ( Fig. 9D View Figure 9 ).

Remarks: There are only slight differences in shell morphology between E. subconsul and E. layanglayang ; the preceding has regular, dense and moderate corrugation at the suture. However, the body colour and colour pattern of E. layanglayang are very different from E. subconsul . Some preliminary evidence from animal morphology and the genetic data suggest that hybridization in the contact zone between E. subconsul and E. layanglayang is occurring: the animal head colour patterns show high consistency within species but the populations in the overlap between the distribution ranges of E. subconsul and E. layanglayang have an intermediate head colour pattern ( Fig. 4G–I View Figure 4 , all from different populations at Crocker Range around 1200 to 1400 m).