Citharexylum, Linnaeus, 1753

Cardoso, Pedro Henrique, Valério, Vanessa Imaculada Dos Reis, Neto, Luiz Menini & Salimena, Fátima Regina Gonçalves, 2021, Verbenaceae in Espírito Santo, Brazil: richness, patterns of geographic distribution and conservation, Phytotaxa 484 (1), pp. 1-43 : 7

3. Dioecious trees; glands in pairs near to the petiole; fruit red.................................................................................... 4. Citharexylum

3’. Monoecious herbs, subshrubs, shrubs or treelets, glands absent near to the petiole; fruit brown, purple or black.

4. Inflorescences compound, fruits divided into 4 units (cluses)