Ganseriella Fedotova et Perkovsky, 2017

Perkovsky, E. E. & Fedotova, Z. A., 2017, The Second Cretaceous Gall Midge Genus Of The Tribe Diallactiini (Diptera, Cecidomyiidae) From The Late Cretaceous Burmese Amber, Vestnik Zoologii 51 (2), pp. 117-124 : 119-123

Genus Ganseriella Fedotova et Perkovsky , gen. n.

urn:lsid:zoobank.org:act:5093FEDA-A88E-485D-ABCD-9DA8145C85A8

T y p e s p e c i e s: Ganseriella pankowskiorum Fedotova et Perkovsky , sp. n.

D e s c r i p t i o n ( figs 1, 1–3; 2, 1 –8; 3, 1–10). Male. Body slightly longer than wings. Head elongate; eyes fused, positioned around head, without narrow bridge, occiput wide, rounded. Antennae 2+14-segmented, scape very long, slightly enlarged distally, basally narrower than round pedicel. All flagellomeres with distinct very narrow and long necks, but basal enlargements have different forms. Basal enlargement on proximal flagellomeres almost parallel-sided, on medial flagellomeres prolonged swollen, on distal flagellomeres slightly swollen or almost round. Basal enlargement of 1st and 2nd flagellomeres slightly narrowed medially. Proximal flagellomeres much wider and longer than distal ones. Basal enlargement of flagellomeres with basal whorls of very short dense setae and medial crenulate whorls of very long setae strongly curved basally. Distal part of basal enlargement with apical whorl of very long setae reaching of next flagellomere. Necks of flagellomeres always longer than basal enlargement, especially in middle flagellomeres. Mouthparts strongly elongate and wide. Palpi much longer than height of head, 4-segmented, the 3rd and 4th ones longest. Wing evenly and very strongly widened medially, vein m+rm slightly curved medially, R 5 slightly curved apically, joining margin of wing behind wing tip; rs very strong; Cu simple, positioned far from margin of wing, reaching wing edge, forming large lobe. Anal vein clear. Cell between veins C and R 1+2 distinctly sclerotized. Costal vein with break beyond its joining point with R 5. Legs completely densely covered with short setae and sparse long setae. Fore and hind femora longer than their tibia, middle femur and tibia. Middle femora shorter than tibia. All tibia subequal in length. Abdomen slightly swollen near middle (2nd–4th segments) and narrowed distally. Abdominal segments covered by very long setae. Genitalia transverse. Gonocoxite wide, slightly rounded on outside margin, with dense long setae and almost rectangular on inner margin. Gonostylus long and strongly enlarged basally, with long apical dent. Tegmen very long, pointed apically, much longer than gonocoxites (lateral view, fig. 3 , 9).

C o m p a r i s o n. The shape of the body, 2+14-segmented antennae, wing venation, long legs and forms of gonocoxites and gonostyles of the new genus imply its close relation to Johnsonomyia Felt, 1908 ( Felt, 1908; Mamaev, 1964, 1966; Spungis, 1985; Jaschhof, Jaschhof, 2013; Jaschhof, 2016), but the new genus differs from it in the elongated segments of palpi, very large eyes, absence of narrow eye bridge, different shape of flagellomeres, very long necks of flagellomeres, presence of very dense long setae of apical whorl of the basal enlargement of flagellomeres, wider and shorter wing, vein R 5 joining the wing margin far behind the wing tip, presence of sclerotisation of the wing cell between veins C and R 1+2; very long pointed tegmen, and smaller body size. The new genus is closely related to Chastomera Skuse, 1889 , which is currently considered a synonym of Haplusia Karsch, 1877 , but differs from it in a longer rs and part of R 1+2 from rs to fusing with C, in the very long palpi, in absence of dorso-apical projection on medial surface of gonocoxite, in the presence of long pointed tegmen, and in the smaller body size. Chastomera Skuse, 1888 stat. resurr. is obviously a separate genus, distinct from Haplusia , as the redescription of the holotype of type species Ch. bella Skuse shows (Skuse, 1888; Kolesik, Gagné, 2016) and does not meet the original diagnosis of Haplusia ( Karsch, 1877) and the numerous diagnoses of the genus Haplusia given earlier ( Mamaev, 1964, 1966; Gagné, 1978; Jaschhof, 2016; Kolesik, Gagné, 2016). The new genus differs from Haplusia in very long pointed tegmen, very wide cell of wing between veins C and R 4+5, and also CuA and wing margin, very long and short dense setae in apical whorl of basal enlargement of flagellomeres (only short in Haplusia ) and very long neck of middle flagellomeres, additionally.in the absence of a dark spot around vein rs and on vein CuA and in the not banded legs.

E t y m o l o g y. The generic name is an arbitrary combination of letters. Gender feminine.

Figures

Fig. 1. Ganseriella pankowskiorum, male, holotype SIZK, Bu–1: 1 — amber with holotype (×3.3); 2 — general appearance (×22); 3 — body and antennae (×29).

Fig. 2. Ganseriella pankowskiorum, male, holotype SIZK, Bu–1: 1 — head and palpi (×189); 2, 3 — genitalia (2 — lateral view, ×422; 3 — ventral view, ×256); 4 — palpi (×247); 5 — flagellomere 5 (×215); 6 — flagellomere 9 (×180); 7 — flagellomere 13 and 14 (×162); 8 — scape, pedicel, flagellomere 1 and 2, head, thorax, wing and base of abdomen (×56). Scale bar 0.1 mm.

Fig. 3. Ganseriella pankowskiorum, male, holotype SIZK, Bu–1 (1–10): 1 — flagellomere 13 and 14; 2 — flagellomere 5; 3 — flagellomere 9; 4 — head, scapus, pedicel and palpi; 5 — wing; 6 — general appearance; 7 — 2nd–5th tarsomeres; 8, 9 — abdominal segments 6–8 and genitalia (8 — view dorsally and 9 — laterally); 10 — thorax.