Amplirhagada sphaeroidea, Köhler, 2010

Köhler, Frank, 2010, Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae), Records of the Australian Museum 62 (2), pp. 217-284 : 263-264

publication ID	https://doi.org/ 10.3853/j.0067-1975.62.2010.1554
publication LSID	lsid:zoobank.org:pub:749A271A-6942-4D4C-B0DF-B968462BFF0C
persistent identifier	https://treatment.plazi.org/id/03F987C5-FFBF-F773-0362-16044926CB5D
treatment provided by	Felipe
scientific name	Amplirhagada sphaeroidea
status	sp. nov.

Treatment

Amplirhagada sphaeroidea View in CoL n.sp.

Type locality ( Fig. 1 View Figure 1 ). Western Australia , Kimberley , Saint George Basin, St. Andrews Island, 15°21'24"S 124°59'46"E; KIS-2–40. Volcanic boulder scree, vine thicket, under rocks (leg. R. Teale, 25 May 2008) GoogleMaps .

Type material. Holotype WAM S34622 View Materials (Pl. 2.10) . Paratypes AMS C463718 (2 preserved specimens, same as holotype), WAM S37029 View Materials (7 preserved specimens, same as holotype),AMS C463719 (shell, 15°21'34"S 125°00'07"E) GoogleMaps , WAM S37027 View Materials (3 preserved specimens, 15°21'34"S 125°00'07"E) GoogleMaps , WAM S37411 View Materials (1 dry shell, 15°21'34"S 125°00'07"E) GoogleMaps .

Additional, non-type material. WAM S37028 View Materials , WAM S37030 View Materials , WAM S37410 View Materials , WAM S37412 View Materials ( St. Andrews Island).

Etymology. From sphaeroeides (Greek = globular, spherical), referring to globular shape of shell.

Sealing strategy. Free sealer. Shell ( Fig. 70A–B View Figure 70 , Pl. 2.10). Globose to semi-globose, with medium high spire; solid to thick. Periphery angulate; upper sector of whorls rather flattened, basal sector rounded. Umbilicus forming a chink, 90–100 percent concealed by columellar reflection. Background colour blends from horn at base to lightly brownish at top of shell; sub-sutural band absent or diffuse, indistinct, brownish; mid-whorl band absent or diffuse, brownish, thin, indistinct, visible on last whorl(s) only; ventral colour cream; outer lip colour differs from shell, dark brownish-purple; inner lip blends from dark pink outside to horn deeper inside. Protoconch c. 2.4 mm in diameter, comprising 1.7 whorls, smooth. Teleoconch smooth, except of faint growth lines. Angle of aperture 45°; outer lip rounded, thick, slightly expanded to expanded, slightly reflected, basal node of lip weak, palatal node absent. Parietal wall of inner lip absent or inconspicuous.

Pallial morphology. Pallial cavity moderately deep, extending± 3 ⁄ 4 whorl; mantle pigmentation mottled, diffuse dark grey. Kidney extending about half of pallial cavity.

Genital morphology ( Figs. 71–72 View Figure 71 View Figure 72 ). Penis straight, more or less of same length as anterior part of oviduct. Vas deferens forms simple loop before entering penis. Penial retractor muscle very short, stubby. Penial sheath evenly thick. Penial verge short (<1 ⁄ 8 penial chamber), slender to spatulate, with pointed tip. Penial wall pustules fused to regular, smooth, diagonal and delicate lamellae on apical portion, becoming more and more oblique towards base of penial chamber. Main stimulatory pilaster well-differentiated; elongated, large, cone-shaped, corrugated, comprising mainly the apical portion of penial chamber and gradually decreasing in size towards base. Three additional, smooth, narrow, rather indistinct pilasters are formed at base of penial chamber. Vas deferens entering penial sheath apically. Vagina rather short, posteriorly inflated; inner vaginal wall with smooth longitudinal pilasters. Spermathecal duct moderately wide, internally with smooth longitudinal pilasters. Spermathecal head globular, connected with oviduct by connective tissue, internally entirely smooth, wall delicate. Free oviduct comprising about half of anterior part of oviduct, or more than half of anterior part of oviduct; more or less straight. Spermoviduct much longer than anterior part of oviduct. Talon embedded in albumen gland anteriorly.

Radular morphology ( Fig. 70D–E View Figure 70 ). Rectangular. Tooth formula C+15+4+18. With 118 rows of teeth, 26.0 rows per mm (n = 1). Central teeth with bluntly pointed, broadly elongate mesocone, length equal to base of teeth; ectocones vestigial. Lateral teeth with bluntly pointed, broadly elongate mesocone, length equal to base of teeth; ectocones small, endocones vestigial. Marginal teeth with broadly elongate mesocones; ectocones smaller and narrower than mesocones; endocones greatly reduced.

Comparative remarks. One of the few species with large, dome-shaped shells. Differs from A. tricenaria and A. regia by dark outer lip and from A. uwinsensis purplish colour of outer lip. Covering of inner penial wall with lamellae, elongate-conical, corrugated main pilaster, and relatively large verge are diagnostic.