Arsapnia Banks, 1897

Murányi, Dávid, Gamboa, Maribet & Orci, Kirill Márk, 2014, Zwicknia gen. n., a new genus for the Capnia bifrons species group, with descriptions of three new species based on morphology, drumming signals and molecular genetics, and a synopsis of the West Palaearctic and Nearctic genera of Capniidae (Plecoptera), Zootaxa 3812 (1), pp. 1-82 : 14

Arsapnia Banks, 1897 stat. rev.

( Fig. 40 )

Arsapnia Banks, 1897 — Banks 1897: 22. (original description, type species Arsapnia decepta Banks, 1897 ).

Capnia Pictet, 1841 — Claassen 1924: 43. (synonymy of Arsapnia Banks, 1897 with Capnia Pictet, 1841 ).

Capnia decepta species group sensu Nelson & Baumann, 1989 — Nelson & Baumann 1989: 310. (definition and revision).

Diagnosis. Male epiproct: B-scl lacking or vestigial; Lb-scl large, fused with Ep-scl; Ep-scl ventrally entire, laterally divided only in the apex, caudal setae present; I-scl and Ec absent. Male Pp: apical part long and wide; Fp long and narrow, divided from Rp. Male Sg: fused with St 9 and Tg 9, vesicle lacking. Female Sg: rounded and narrow; small lateral sclerites present. Male tergites: Tg 7 with process. Ventral thoracic sclerites: MPrs and MeFs triangular, MeFsp separated from MePfs. Macropterous wings: forewing A1 beyond a and R1 before r curved.

Species included. Type species: Arsapnia decepta Banks, 1897 comb. rev.; Further species: Arsapnia arapahoe ( Nelson & Kondratieff, 1988) comb. n., Arsapnia coyote ( Nelson & Baumann, 1987) comb.n., Arsapnia pileata ( Jewett, 1966) comb. n., Arsapnia sequoia ( Nelson & Baumann, 1987) comb.n., Arsapnia teresa ( Claassen, 1924) comb. n., Arsapnia tumida ( Claassen, 1924) comb. n., Arsapnia utahensis ( Gaufin & Jewett, 1962) comb. n. All these West Nearctic species were examined (see Appendix 1).

Remarks. Banks (1897) erected the genus on the basis of several wing venation characters. Claassen (1924) synonymised it with Capnia on the basis of Pictet’s (1841) drawings and description of the type species, C. nigra . Indeed, the venation shows much similarity between the two genera, but male terminalia, especially the epiproct differ greatly and there seems to be no reason to retain the genus Arsapnia in synonymy with Capnia . Herein, the genus is proposed only for members of the C. decepta group sensu Nelson & Baumann 1989, however, the C. s.l. excavata and C. s.l. nana groups sensu Nelson & Baumann 1989 may also belong to this genus. Nevertheless, we exclude C. s.l. nelsoni Kondratieff & Baumann, 2002 , because of its different genitalic structures (see Capnia sensu lato).

Figures

FIGURES 32–48. Male terminalia of Capniidae, ventral view—32: Zwicknia bifrons (Newman); 33: Bolshecapnia gregsoni (Ricker); 34: Eucapnopsis stigmatica transversa Aubert; 35: Isocapnia grandis (Banks); 36: Capnia s.l. arensi Zhiltzova; 37: C. s.l. prolongata; 38: C. s.l. pedestris Kimmins; 39: C. s.s. nigra (Pictet); 40: Arsapnia decepta Banks; 41: Allocapnia granulata (Claassen); 42: Mesocapnia silvatica Raušer; 43: Capnura venosa Banks; 44: Utacapnia lemoniana (Nebeker & Gaufin); 45: Paracapnia opis (Newman); 46: Nemocapnia carolina Banks; 47: Capnopsis schilleri schilleri (Rostock); 48: Capnioneura nemuroides Ris (C: cercus; Ep-scl: main epiproct sclerite; Fp: fusion plate; Pp: paraproct; Sg: subgenital plate; St 9: sternite 9; Tg 9: tergite 9; Tg 10: tergite 10; Vv: ventral vesicle)—scale 1 mm.