Heteropoda simplex Jäger and Ono, 2000

Bayer, Steffen & Jäger, Peter, 2009, Heteropoda species from limestone caves in Laos (Araneae: Sparassidae: Heteropodinae), Zootaxa 2143 (1), pp. 1-23 : 15-20

publication ID

https://doi.org/ 10.11646/zootaxa.2143.1.1

persistent identifier

https://treatment.plazi.org/id/03F9C605-2B5A-FFDD-1F8B-FD24985DFF39

treatment provided by

Felipe

scientific name

Heteropoda simplex Jäger and Ono, 2000
status

 

Heteropoda simplex Jäger and Ono, 2000

Figs 73–111 View FIGURES 73–85 View FIGURES 86–95 View FIGURES 96–103 View FIGURES 104–111 , 115 View FIGURES 112–117

Heteropoda simplex Jäger and Ono 2000: 49 , figs 25–26 (description of ♂).

Heteropoda venatoria (Linnaeus) (ad part).— Jäger and Ono (2000: 47, only ♀♀, figs 20–21; misidentification).

Type material: Holotype ( PJ 1298 ) , ♁, JAPAN: Okinawa Prefecture: Iriomotejima Island , 29 July 1987, A. Tanikawa ( NSMT) . Paratype: 1 ♂ ( PJ 1296 ) , same data as for holotype, March 1986, A. Tanikawa ( NSMT) . All examined .

Additional material examined (8 ♁, 28 ♀♀). JAPAN: Okinawa Prefecture: Iriomotejima Island : 1 ♀ ( PJ 1313 ) , Otomi , 4 January 1989, A. Tanikawa ( NSMT) . 1 ♀ ( PJ 1325 ) , Otomi , 31 March 1987, A. Tanikawa ( NSMT) . 1 ♂ ( PJ 1500 ) , 17 December 1965, T.A. Vehida ( NSMT) . 1 ♀ with egg sac ( PJ 1530 ) , Mt. Bannadake , 22 October 1990, A. Yawata ( NSMT) . 1 ♀ ( PJ 3011 ) , 29 July 1985, A. Tanikawa ( NSMT) . 1 ♂ ( PJ 3016 ) , Ohara , 20 June 2008, A. Tanikawa ( SMF 58955 View Materials ) . LAOS: Luang Nam Tha Province: Vieng Phoukha : 1 ♀ ( PJ 2789 ) , Tham Pasat Thia 1 (Map F 47-119/002), N 20°46'44.1", E 10°00'51", sample 077/06, 09 February 2006, H. Steiner ( SMF 58950 View Materials ) GoogleMaps . 1 ♀ ( PJ 2845 ) , Tham Pasat , NW-entrance, resurgence cave, N 20°46'37.2'', E 101°01'00.2'', 705 m, cave below, 4 March 2008, P. Jäger by hand ( SMF 58953 View Materials ) GoogleMaps . 1 ♀ ( PJ 3086 ) , Tham Pasat, Tham Nam Mai , N-entrance, inside cave, N 20°46'37.2'', E 101°01'00.2'', 705 m, 5 March 2008, P. Jäger by hand,( SMF 58961 View Materials ) GoogleMaps . 1 ♁ ( PJ 2838 ) , Tham Nam Eng , N 20°43'26.8'', E 101°09'00.3'', 739 m, secret cave, 5 March 2008, P. Jäger by hand ( SMF 58951 View Materials ) GoogleMaps . 1 ♀ ( PJ 3089 ) , same data as for preceding specimen ( NSMT) . 1 ♀ ( PJ 2839 ) , same data as for preceding specimen ( SMF 58952 View Materials ) . 1 juv. (SD 570), same data as for preceding specimen ( SMF) . LAOS: Luang Prabang Province: 1 ♀ ( PJ 2468 ) , Nam Ou, Nong Khiao, Tham Pathok , N 20°33.082', E 102°37.925', 373 m, inside cave, 10 March 2006, P. Jäger & J. Altmann by hand at night ( SMF 58963 View Materials ) GoogleMaps . 1 ♀ ( PJ 2470 ) , same locality as for preceding specimen, SD 512, 11 March 2006, P. Jäger & J. Altmann ( SMF 59095 View Materials ) . 1 ♀ ( PJ 2469 ) , same data as for preceding specimen, SD 515 ( SMF 59094 View Materials ) . 1 ♀ ( PJ 2544 ) , same locailty as for preceding specimens, 26 March 2007, P. Jäger & F. Steinmetz ( SMF 58967 View Materials ) . 1 ♀ ( PJ 3110 ) , same data as for preceding specimens, 27 March 2007 ( SMF 58968 View Materials ) . 1 ♀ ( PJ 2546 ) , same data as for preceding specimens, 16–17 March 2007 ( SMF 58964 View Materials ) . 1 ♀ ( PJ 2756 ) , same data as for preceding specimens ( SMF 58966 View Materials ) . 1 ♀ ( PJ 2714 ) , same data as for preceding specimens ( SMF 58965 View Materials ) . 1 ♀ ( PJ 2549 ) , same locality as for preceding specimens, outside cave, banana fields, 18 March 2007, P. Jäger & F. Steinmetz by hand ( SMF 58945 View Materials ) . 1 ♀ ( PJ 2745 ) , same locality as for preceding specimen, inside limestone cave, 24 March 2007, F. Steinmetz by hand ( MHNG) . 1 ♂ ( PJ 2768 ) , same locality as for preceding specimens, 5 m outside 3. cave, 27 March 2007, F. Steinmetz by hand ( SMF 58946 View Materials ) . 1 ♂ ( PJ 3012 ) , same data as for preceding specimen, outside cave, leaflitter, 29 March 2007, P. Jäger & F. Steinmetz by hand at night ( MHNG) . 1 ♂ ( PJ 3019 ) , same locality as for preceding specimen, outside cave, at stone, 23 March 2007, F. Steinmetz ( SMF 58956 View Materials ) . 1 ♀ ( PJ 2849 ) , SE Luang Prabang, Nam Khan, Ban Keng Koung , 372 m, N 19°40.963', E 102°18.442', waterfall, on rock, 15 February 2008, P. Jäger ( SMF 58954 View Materials ) GoogleMaps . 1 ♀ ( PJ 3084 ) , same data as for preceding specimen ( AMNH) . 1 ♀ ( PJ 3049 ) , Tham Pha (Map E 48-01/11), N 19°45'43.1'', E 102°01'42.6'', sample 038/04, LP29-1, entrance hall, 29 December 2003, H. Steiner ( SMF 58958 View Materials ) GoogleMaps . LAOS: Huaphan Province : 1 ♂ ( PJ 2784 ) , Xam Neua, Tham Nam (Map F 48-125-008), N 20°27'15.0", E 104°10'4.7", sample 124/07-1, 15 January 2007, H. Steiner ( SMF 58947 View Materials ) GoogleMaps . 1 ♀ ( PJ 2785 ) , same data as for preceding specimen, sample 124/07-2 ( SMF 58948 View Materials ) . 1 ♀ ( PJ 2786 ) , same data as for preceding specimens, sample 124/07-3 ( SMF 58949 View Materials ) . 1 ♀ ( PJ 3065 ) , same data as for preceding specimens, sample 080/08, 18 January 2008 ( SMF 58959 View Materials ) . 1 ♂ ( PJ 3048 ) , Tham Kouang Tai (Map F 48-125-020), N 20°27'28.2", E 104°09'51.0'', sample 097/08, 12 January 2008, H. Steiner ( SMF 58957 View Materials ) GoogleMaps . 1 ♀ ( PJ 3066 ) , same data as for preceding specimen ( MNHN) . 1 ♀ ( PJ 3070 ) , Tham Sang Lek Hok (Map F 48-125-036), N 20°28'03.6", E 104°10'42.5'', sample 075/08, 078/08, 11 January 2008, H. Steiner ( SMF 58960 View Materials ) GoogleMaps . 2 ♀ (PJ 3090–3091), Vieng Thong, Tham Nam Lot (Map F 48-135-004), N 20°14'56.2'', E 103°23'47'', sample 31/09, Northern Lao- European Cave Project 2009, SD 593 ( PJ 3090 ) GoogleMaps , 13 January 2009, H. Steiner, ( SMF 59011 View Materials ) .

Extended diagnosis. Tibia, cymbium and tegulum of male palp not distinctly elongated (cymbium less that 2.6 times as long as tegulum) ( Fig. 74 View FIGURES 73–85 ), dRTA of male palp narrow and shorter than in H. aemulans sp. nov. ( Fig. 77 View FIGURES 73–85 ); vRTA with distinct small hump ( Fig. 75 View FIGURES 73–85 ). Sperm duct distinctly S-shaped ( Figs 76 View FIGURES 73–85 , 86–90 View FIGURES 86–95 ). Septum of epigyne freely visible, just slightly covered by lateral lobes, mainly in the posterior part. Anterior part of septum between copulatory openings more (or far more) than half as broad as its medial, broadest part ( Fig. 85 View FIGURES 73–85 ). Anterior part of internal duct system longer than wide (in some specimens only slightly longer), median part of first winding diverging anteriorly and posteriorly ( Fig. 82 View FIGURES 73–85 , 104–111 View FIGURES 104–111 ); posterior part of internal duct system not or just slightly covered by anterior part ( Fig. 82 View FIGURES 73–85 ).

Redescription. Male (n=10):

Prosoma length 6.6 – 11.8, prosoma width 5.9 – 11.1, anterior width of prosoma 2.8 – 5.2, opisthosoma length 8.5 – 12.1, opisthosoma width 4.7 – 6.2. Eyes: AME 0.32 – 0.44, ALE 0.53 – 0.76, PME 0.39 – 0.61, PLE 0.55 – 0.77, AME – AME 0.17 – 0.38, AME – ALE 0.04 – 0.13, PME – PME 0.26 – 0.52, PME – PLE 0.40 – 0.76, AME – PME 0.38 – 0.67, ALE – PLE 0.43 – 0.76, clypeus height at AME 0.65 – 1.16, clypeus height at ALE 0.38 – 0.85.

Cheliceral furrow with denticles situated close to three promarginal teeth and further distally ( Fig. 80 View FIGURES 73–85 ), in most specimen even over entire furrow. Retrolateral margin with 4, rarely 5 (n=1) teeth ( Figs 79–80 View FIGURES 73–85 ).

Spination: Palp: 131, 101, 2121; legs: femur I–II 323, III 333(332, n=2), IV 331(332, n=1); patella I – III 101, IV 101(001, n=4); tibia I – II 2226, III 2226 (2126, n=3), IV 2126 (2226, n=2); metatarsus I 1014 (0014, n=1), II 1014 (2014, n=1), III 2014, IV 3036. Leg formula: 2143. Measurements of palp and legs: Palp 10.3 – 17.1 [3.4 – 5.9, 1.5 – 2.6, 2.4 – 3.8, 3.0 – 4.8], I 36.6 – 62.3 [9.8 – 16.6, 3.6 – 6.7, 10.5 – 18.2, 9.8 – 18.4, 2.9 – 4.5], II 40.9 – 70.4 [11.9 – 18.8, 4.0 – 7.0, 11.6 – 20.7, 10.4 – 20.3, 3.0 – 4.6], III 31.1 – 55.0 [9.3 – 15.7, 3.1 – 5.9, 8.7 – 15.6, 7.6 – 14.9, 2.4 – 3.5], IV 34.1–59.5 [9.8 – 16.5, 3.0 – 5.5, 9.3 – 16.5, 9.3 – 17.5, 2.7 – 4.0].

Pedipalp as in diagnosis. Conductor relatively long and sabre-like, arising in a 9- to 10-o’clock-position on tegulum ( Figs 74, 76 View FIGURES 73–85 ). Ventral flange of conductor turning prolaterally basally, before the basal quarter of conductor ( Fig. 76 View FIGURES 73–85 ). Embolus arising in a 6-o’clock-position on tegulum, running a semi-circle. Male palp with RTA arising distally on tibia ( Figs 74–75 View FIGURES 73–85 ).

Colouration. Generally yellowish-brown with darker brown pattern on opisthosoma, with darker brown annulate patches on legs, and with dark brown radial patches on prosoma. Light interspaces between darker annulate patches and other large patches on legs, especially on femora, containing several small, darker brown spots (♀, Fig. 115 View FIGURES 112–117 ).

Description. Female (n=19):

Prosoma length 6.4 – 13.2, prosoma width 5.9 – 11.3, anterior width of prosoma 3.3 – 5.8, opisthosoma length 9.4 – 17.5, opisthosoma width 5.3 – 11.1. Eyes: AME 0.32 – 0.56, ALE 0.51 – 0.82, PME 0.40 – 0.63, PLE 0.54 – 0.79, AME – AME 0.18 – 0.38, AME – ALE 0.07 – 0.17, PME – PME 0.29 – 0.55, PME – PLE 0.40 – 0.88,

AME – PME 0.44 – 0.70, ALE – PLE 0.52 – 0.85, clypeus height at AME 0.67 – 1.40, clypeus height at ALE 0.46 – 1.24.

Cheliceral furrow with denticles distributed in most specimens over entire furrow, if not then at least to the subdistal tooth of retromargin ( Fig. 79 View FIGURES 73–85 ). 3 teeth promarginally and 4 (rarely 5, n=2) retromarginally.

Palpal claw ( Fig. 78 View FIGURES 73–85 ) with 8 (n=10) or 7 (n=6), rarely 9 (n=3) teeth. Arising points of the teeth forming a curved line ( Fig. 78 View FIGURES 73–85 ).

Spination: Palp: 131, 101, 2121, 1014; legs: femur I 323, II 323(333, n=1), III 333(332, n=3), IV 331(332, n=1); patella I – IV highly variable (101, 100, 000, 001); tibia I 2026, II 2026 (2126, n=2), III 2126 (2226, n=2, 2026, n=1), IV 2126 (2026, n=2); metatarsus I – II 1014, III 2014, IV 3036. Leg formula: 2143(2413, n=1). Measurements of palp and legs: Palp 11.8 – 18.6 [2.9 – 5.9, 1.5 – 3.0, 2.7 – 5.2, 3.0 – 5.8], I 25.3 – 60.1 [7.3 – 16.1, 3.2 – 6.8, 10.3 – 17.4, 9.1 – 15.0, 2.6 – 3.8], II 41.6 – 64.3 [11.5 – 17.9, 4.5 – 6.7, 10.9 – 18.5, 9.9 – 17.1, 2.7 – 4.1], III 33.0 – 49.7 [10.1 – 15.5, 3.5 – 6.2, 9.2 – 15.3, 7.9 – 14.1, 2.3 – 3.5], IV 36.5 – 59.6 [10.6 – 16.3, 3.5 – 5.8, 9.9 – 16.5, 9.9 – 17.0, 2.6 – 4.0].

Copulatory organ as in diagnosis. Epigynal field with long anterior bands (length/basal length 1.38–1.65). Septal pocket of epigyne mostly elongated triangular ( Fig. 85 View FIGURES 73–85 ). Slit sense organs mostly outside of epigynal field, which latter is almost quadratic or stout rectangular. Anterior part of internal duct system longer than wide (in a few specimens only slightly longer) and with more than one winding until turning point ( Figs 82–83 View FIGURES 73–85 ); posterior part in most cases more or less round in shape ( Fig. 82 View FIGURES 73–85 ).

Colouration. As described in male ( Fig. 115 View FIGURES 112–117 ).

Variation of copulatory organs. Despite the very large distribution range of this species, male pedipalp varies barely in shape ( Figs 86–90 View FIGURES 86–95 ). Distinct differences were observed in palp size ( Figs 86–95 View FIGURES 86–95 ; as known from H. tetrica: Eusemann & Jäger in print). Anterior bands of epigynal field vary in length, specimens from Japan and from the eastern part of Laos (Huaphan Province) seem to have longer anterior bands ( Figs 96, 103 View FIGURES 96–103 ). In some specimens grooves appearing on median septum of epigyne ( Fig. 102 View FIGURES 96–103 ). Few specimens with septal pocket relatively short ( Fig. 98 View FIGURES 96–103 ). Helical sections of internal duct system, and posterior part varying in size and shape ( Figs 104–111 View FIGURES 104–111 ). These different characteristics do not seem to be congruent with geographical distribution. Distance between median part of first winding differing ( Figs 108 and 110 View FIGURES 104–111 ). In rare cases anterior part covering about 1/5 of the length of posterior part ( Fig. 105 View FIGURES 104–111 ), but not as distinct as in H. maxima or H. steineri sp. nov. (≥ 1/3 covering).

NSMT

National Science Museum (Natural History)

SMF

Forschungsinstitut und Natur-Museum Senckenberg

MHNG

Museum d'Histoire Naturelle

AMNH

American Museum of Natural History

MNHN

Museum National d'Histoire Naturelle

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Araneae

Family

Sparassidae

Genus

Heteropoda

Loc

Heteropoda simplex Jäger and Ono, 2000

Bayer, Steffen & Jäger, Peter 2009
2009
Loc

Heteropoda simplex Jäger and Ono 2000: 49

Jager, P. & Ono, H. 2000: 49
2000
Loc

Heteropoda venatoria (Linnaeus)

Jager, P. & Ono, H. 2000: 47
2000