Cephennium fractum, Assing & Meybohm, 2021

Assing, Volker & Meybohm, Heinrich, 2021, The Cephennium fauna of Turkey and the Middle East (Coleoptera: Staphylinidae: Scydmaeninae), Beiträge Zur Entomologie = Contributions to Entomology 71 (1), pp. 1-28 : 8

Cephennium fractum spec. nov.

urn:lsid:zoobank.org:act:26CE8301-2927-4893-B93E-9F2BEF4F7F07

( Figs 2–3 , 32 , 46 , Map 2 )

Type material examined: Holotype : “ N39°50'08 E28°25'31 (19), TR Bursa 16.4.2010, Devecikonagi 640 m, leg. Brachat & Meybohm / Holotypus  Cephennium fractum sp. n., det. V. Assing 2020” (cAss) . Paratypes: 2 , 2 : same data as holotype (cAss); 1 , 2 : “ N40°06'47 E029°04'10 (21), TR Bursa 17.4.2010, Uludag 1430 m, leg. Brachat & Meybohm” (cAss); 4 : “ N40°31'40 E029°52'10 (27), TR Bursa 20.4.2010, Samanli Daglari 740 m, leg. Brachat & Meybohm” (cAss); 1 : “ N40°30'41 E029°49'50 (26), TR Bursa 19.4.2010, Samanli Daglari 720 m, leg. Brachat & Meybohm” (cAss); 1 , 1 : “TR – Balikesir (11), Kaz Daği-Gebiet, ca. 20 km NO Edremit, Kuruçam Tepe , 708 m, 13.IV.2010 / N39°41'39,2 E27°08'31,7, leg. Meybohm & Brachat” (cAss); 2 , 2 : “ N40°38'47 E30°03'25 (2), TR Kocaeli 22.4.2014, Pazarcayiri 880 m, Brachat & Meybohm” (cAss); 1 , 1 : “ N40°37'43 E30°03'36, TR Kocaeli 25.4.2012, Pazarcayiri 720 m, Brachat & Meybohm (5)” (cAss); 1 : “ N40°38'47 E30°03'25, TR Kocaeli 25.4.2012, Pazarcayiri 880 m, Brachat & Meybohm (6)” (cAss); 1 , 2 : “ N40°39'04 E30°03'42, TR Kocaeli 24.4.2012, Kartepe 1030 m, Brachat & Meybohm (1)” (cAss); 1 , 3 : “ N40°38'54 E30°05'32, TR Kocaeli 5.5.2012, Kartepe 1270 m, Brachat & Meybohm (35)” (cAss); 1 , 2 : “ N40°39'24 E30°04'46, TR Kocaeli 24.4.2012, Kartepe 1140 m, Brachat & Meybohm (2)” (cAss); 1 : “ N39°42'45 E27°10'23 (14), TR Canakkale, 14.4.2010, Kurucam Tepe 600 m, leg. Brachat & Meybohm” (cAss) .

Etymology: The specific epithet is the past participle of the Latin verb frangere (to fold, to break) and alludes to the subapically distinctly curved male protibiae.

Description: Body length 1.1–1.3 mm; length of pronotum 0.38–0.42 mm; width of pronotum 0.45–0.51 mm; length of antennae 0.55–0.60 mm. Habitus as in Figs 2–3 . Colouration: body reddish to brown, often with the elytra somewhat darker than the head and pronotum.

Eyes composed of 6–8 ommatidia. Antenna of similar shape as in C. bodemeyeri .

Pronotum approximately 1.2 times as broad as long, broadest in anterior half, and moderately convex in crosssection; lateral margins straight in posterior half.

Elytra noticeably broader than pronotum, anteriorly with a subcircular impression with tomentose whitish pubescence; humeral carina fine, approximately as long as the combined length of the basal three antennomeres.

: protibia distinctly bent subapically and flattened in subapical portion, apically dilated; metaventrite extensively flattened, but not distinctly impressed; aedeagus ( Figs 32 , 46 ) long and slender, 0.60–0.66 mm long; ventral process constricted in the middle and apically with a moderately long apical extension in ventral view; internal sac with weakly sclerotized small structures; paramere long and straight, extending to apical fifth of ventral process, with a long and weakly curved apical seta.

Comparative notes: This species is distinguished from the similar C. bodemeyeri by slightly shorter antennae, slightly smaller eyes, smaller and more circular anterior tomentose impressions on the elytra, subapically more sharply bent male protibiae, and a longer and more slen- der aedeagus with a differently shaped ventral process with a shorter apical extension, by straight parameres, and by internal structures of different shapes.

Distribution and natural history: The distribution is confined to Bursa, Balıkesir, Kocaeli, and Çanakkale provinces, Northwest Anatolia ( Map 2 ). The specimens were sifted from litter in beech and hornbeam forests, mostly with rhododendron undergrowth and often in dark stream valleys. The altitudes range from 600 to 1430 m. In one locality in Kocaeli, the species was collected together with C. bodemeyeri .

Figures

Map 2: Distributions of Cephennium fractum (black circles) and of the species of the C. ericinum group: C. digitale (white triangle); C. spinigerum (black triangle); C. victrix (white circles); C. varispinosum (black diamond); C. ericinum (white diamond).

Figs 1–8: Male habitus in dorsal view (1–6) and head and pronotum in lateral view (7–8) of Cephennium bodemeyeri (1), C. fractum (2–3), C. infractum (4), C. immutatum (5), C. ericinum (6–7), and C. varispinosum (8). Scale bars: 1–6: 1.0 mm; 7–8: 0.5 mm.

Figs 25–35: Aedeagus in ventral view (25, 27, 29–30, 32–35) and apical portion of aedeagus in ventral view (26, 28, 31) of Cephennium perispinctum (25–26), C. fibuliferum (27–28), C. bodemeyeri (29–31), C. fractum (32), C. infractum (33), and C. immutatum (34–35). Scale bars: 25, 27, 29–30, 32–35: 0.2 mm; 26, 28, 31: 0.1 mm.

Figs 36–47: Aedeagus in ventral view (36, 38, 40, 42, 44), internal structures of aedeagus (37, 39, 41, 43, 45), and apical portion of aedeagus in ventral view (46–47) of Cephennium ericinum (36–37), C. varispinosum (38–39), C. victrix (40–41), C. spinigerum (42–43), C. digitale (44–45), C. fractum (46), and C. infractum (47). Scale bars: 36, 38, 40, 42, 44: 0.2 mm; 37, 39, 41, 43, 45–47: 0.1 mm.