Procryphalus petioli (Beaver) Beaver & Smith & Sanguansub, 2019

Beaver, Roger A., Smith, Sarah M. & Sanguansub, Sunisa, 2019, A review of the genus Dryocoetiops Schedl, with new species, new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae), Zootaxa 4712 (2), pp. 236-250 : 247-248

Procryphalus petioli (Beaver) comb. n.

Dryocoetiops petioli Beaver 1990: 281 .

Taxonomy. We have examined the type series (NHML, RAB) and further specimens collected by RAB at the type locality. The species was provisionally placed in the genus Dryocoetiops on the suggestion of S. L. Wood who ex- amined some specimens from the original collection ( Beaver 1990). However, dissections have shown that males are present, apparently indistinguishable from females using external morphology, and there is probably a 1: 1 sex ratio, unlike Dryocoetiops . B. Jordal (personal communication, January 2015) has found that the male genitalia are of the cryphaline type. Jordal and Cognato (2012), and Gohli et al. (2017) have shown, based on multiple genetic loci, that the species is closely related to two species of the cryphaline genus, Procryphalus Hopkins, 1915 ( P. fraxini (Berger, 1916) , P. mucronatus (Leconte, 1879)) . The species is morphologically very distinct from these species, but it is provisionally transferred here to Procryphalus .

Distribution. Thailand.

Biology. Collected only from the petioles of fallen leaves of Manglietia garrettii Craib (Magnoliaceae) , incorrectly given as M. garelli in Beaver (1990) . The beetles bored into the base of the petiole. The species appears to be monogamous. Immature stages were not found in the galleries, and it is possible that leafstalks are used primarily as a ‘refuge’ in the absence of other suitable breeding material.