Anthomyza pengellyi, Roháćek & Barber, 2016

Roháćek, Jindřich & Barber, Kevin N., 2016, Nearctic Anthomyzidae: a monograph of Anthomyza and allied genera (Diptera), Acta Entomologica Musei Nationalis Pragae (suppl.) 56, pp. 1-412 : 144-161

Anthomyza pengellyi sp. nov.

( Figs 256, 257 , 260–274 , 294 )

Type material. HOLOTYPE: ♂, “CAN:AB: Elbow-Sheep Wi-ldland Pk , ~22.5kmNW Hi-ghwood House , 14.vii.2011, KNBarber, sweeps, gras-ses incl. Poa compressa 50°31.38’N 114°53.17’W ”, “debu 01502616” and “ Holotypus ♂ Anthomyza pegellyi sp. n., J. Roháček & K. N. Barber det. 2014” (red). The specimen is in perfect condition (see Fig. 294 ), with well visible gonostyli ( DEBU, intact). PARATYPES: CANADA: ALBERTA: Banff, Norquay Mt. Meadows, 5000–6000', 4.vi.1925, 1 ♀, O. Bryant leg. ( USNM); Banff N. P., 4500', 28.vii.1967, 1 ♀, J. R. Vockeroth leg.; Banff N. P., Eisenhower Jct., 4700', 6.vii.1955, 1 ♀, J. R. McGillis leg.; Banff [N. P.], Eisenhower Jct., 4700', Malaise trap, 9.vii.1962, 1 ♀, Mason leg. (all CNCI); Banff N. P., Mt. Eisenhower, 27.vii.1967, 1 ♂, B. A. Foote leg. ( USNM); ~21.9 km W Bearberry, Hwy #734, 51°56.18'N 115°11.76'W, sweeps, wet roadside, Equisetum ssuviatile , 24.vii.2008, 1 ♂, sweeps, mostly Calamagrostis canadensis , 24.vii.2008, 1 ♂; ~22.7 km S Bellevue, Hwy 774, 49°22.62'N 114°22.58'W, sweeps, roadside ditch, mostly Carex spp., Equisetum , grasses, 17.vii.2011, 1 ♂, all K. N. Barber leg. (all CNCI); Bilby, 1.vi.1924, 1 ♀, 22.vi.1924, 1 ♂ 1 ♀, 12.vii.1924, 1 ♂, O. Bryant leg. ( USNM); Hwy 97, Blue Lake, 34 km NW Williams Lake, 23.vi.1978, 1 ♂, P. H. Arnaud Jr. leg. ( CASC); Cadomin, base of Prospect Mt., mouse runs in grass, 18.vii.1987, 1 ♂ 2 ♀♀, S. A. Marshall leg. ( DEBU); ~ 1.2 km S Cadomin, entrance to Whitehorse Wildland P. Pk., 53°00.70'N 117°20.05'W, sweeps, Bromus inermis , Hedysarum boreale , 23.vii.2008, 5 ♂♂ 5 ♀♀; ~ 3.7 km NNE Cadomin, Hwy #40, 53°03.84'N 117°17.82'W, sweeps, roadside Bromus inermis , 23.vii.2008, 1 ♂ 1 ♀, all K. N.Barber leg. (all CNCI); Calgary,Fish Creek P. Pk., 50°55.581'N 114°07.399'W, sweep by river, 11.vi.2010, 1 ♀; same locality but 50°54.407'N 114°01.119'W, sweep brush by ranch house, 25. vi.2010, 1 ♂, both J. E. Swann leg. (both BDUC); Cooking Lk., 13.vi.1937, 1 ♂ 1 ♀, F. O. Morrison leg. ( LEMQ 0039947, -65); Dunvegan, 55°55.39'N 118°35.74'W, sweep south facing grass slope at dusk, 19.vii.2003, 7 ♂♂ 1 ♀, S. Boucher leg. ( LEMQ 0040436, -37, -39, -41–45); Dunvegan, N shore of Peace River, sweep south facing grass slope at dusk, 12.vii.1997, 6 ♂♂ 2 ♀♀ ( LEMQ 0039944, -45, -57–60, -62, -63); Dunvegan, n. shore of Peace River, sweep grasses at edge of agricultural neld, 13.vii.1997, 8 ♂♂ 5 ♀♀ ( LEMQ 0039912, -23, -26, -32, -43, -48, -88–92, -98, -99), sweep grasses along agricultural neld, 14.vii.1997, 1 ♂ 3 ♀♀ ( LEMQ 0039993, -0040000, -01, -07), all T. A. Wheeler leg.; Dunvegan, N side Peace River, pan traps in grasses, 22.v.–6.vi.1995, 1 ♀ ( LEMQ 0039909), 6–19.vi.1995, 1 ♂ ( LEMQ 0039908), S. Boucher leg.; Dutch Creek Bridge, Hwy 940, 3.vii.1982, 1 ♀, B. V. Peterson leg. ( CNCI); Edmonton, 11.vi.1937, 1 ♂, E. H. Strickland leg. ( LEMQ); Edmonton, 14.vi.1966, 2 ♂♂ 1 ♀, K. A. Spencer leg. ( BMNH, 1 ♂ 1 ♀ genit. prep.); Edmonton, prairie, 12.vi.1937, 1 ♀, E. H. Strickland leg. ( USNM); Edmonton, Ft. Edmonton Pk., 53°30.3'N 113°34.2'W, sweeps, veg. banks of N. Sask[atchewan] R., 14.vii.1999, 2 ♀♀, K.N.Barber leg.( CNCI); Elbow-Sheep Wildland Pk., ~22.5 km NW,Highwood House, 50°31.38'N 114°53.17'W, sweeps, Carex utriculata , 14.vii.2011, 5 ♂♂ 6 ♀♀ ( DEBU 01502573–83), sweeps, grasses incl. Poa compressa , 14.vii.2011, 21 ♂♂ 14 ♀♀ ( DEBU 01502609–15, -17–44), pooter, Carex utriculata , 14.vii.2011, 1 ♀ ( DEBU 01502594), pooter, Carex utriculata , 16.vii.2011, 2 ♂♂ 3 ♀♀ ( DEBU 01503655–59); ~ 22.5 km NW Highwood House, ~ 4 km W Mist Ck., 50°31.38'N 114°53.17'W, sweeps, Carex sp. (large), 25.vii.2008, 2 ♂♂, sweeps, Carex sp. (small), 25.vii.2008, 1 ♀ ( DEBU); Elbow-Sheep Wildland Pk., ~ 22.5 km NW Highwood House, 50°31.40'N 114°53.12'W, sweeps, Carex utriculata , 14.vii.2011, 1 ♂ ( DEBU 01502661); same locality but ~ 26 km NW Highwood House, Mt. Lipsett day use, 50°33.26'N 114°57.27'W, sweeps, various grasses, 14.vii.2011, 1 ♂ ( DEBU 01502568); ~ 26 km NW Highwood House, Mt. Lipsett pulloff, 50°33.26'N 114°57.27'W, sweeps, Bromus inermis , Leymus innovatus , 25.vii.2008, 10 ♂♂ 16 ♀♀ ( DEBU), all K. N. Barber leg.; Elkwater, 5.vi.1952, 1 ♂, 8.vi.1952, 3 ♀♀, A. R. Brooks leg., 8.vi.1952, 1 ♀, L. A. Konotopetz leg.; Elkwater Park, 26.v.1952, 1 ♂, A. R. Brooks leg., 31.v.1952, 1 ♀, 12.vi.1952, 1 ♂, L.A. Konotopetz leg. (all CNCI); 4 km SE Grimshaw, sweep grasses at margin of wheat neld, 13.vii.1997, 1 ♂ 1 ♀ ( LEMQ 0039913, 0040010), sweep vegetation at edge of wheat neld, 13.vii.1997, 2 ♂♂ ( LEMQ 0039966, -67), S. Boucher leg., sweep margin of agricultural neld, 13.vii.1997, 2 ♂♂ ( LEMQ 0039970, -40002), T. A. Wheeler leg.; [Fort] McMurray, 3.vii.1953, 1 ♀, 10.vii.1953, 5 ♂♂ 7 ♀♀, 4.viii.1953, 5 ♂♂ 1 ♀, 7.viii.1953, 2 ♀♀, 8.viii.1953, 1 ♂ 2 ♀♀, 10.viii.1953, 2 ♂♂; nr. [Fort] McMurray, 29.v.1953, 1 ♂, all G. E. Ball leg. (all CNCI); 2 km N of Fortree Mountain Ski Resort Road jct. Hwy 40, 1700 m, Alta 6 meadow, 21.vii.1986, 1 ♂, H. Goulet leg.; Halfway Slough, 59°56.049'N 111°44.344'W, sweep, 5.vii.2012, 1 ♂, S. & K. Williamson leg. (both BDUC); ~ 3.4 km SSW Hinton, Hwy #40, 53°21.24'N 117°37.35'W,sweeps,mostly Bromus inermis , 23.vii.2008, 1 ♂; ~ 11 km WSW Hinton, nr. jct. Twp. Rd. 510A & Range Rd. 262, 53°22.72'N 117°44.13'W, sweeps, mostly Bromus inermis , 22.vii.2008, 3 ♂♂ 2 ♀♀, all K. N. Barber leg. (all CNCI); Jasper, 16.vi.1966, 2 ♂♂, K.A. Spencer leg. ( BMNH); 2 mi S Jasper, 30.vii.1967, 1 ♂, S. P. Whitney leg. ( USNM); Kananaskis, 0.3 km N of Barrier Lk. Dam, 51°02'N 115°03'W, sweep, 2.viii.2001, 1 ♀, C. J. Borkent leg. ( RBCM ENT 006-007073); Kananaskis, Mt. Loretto [sic Lorette] Pond[s], 50°38.076'N 113°06.649'W, Bioblitz, sweep, 16.vii.2011, 1 ♂, G. Shrestha & E. Nelson leg. ( BDUC); Kananaskis Country, nr. Beehive Natural Area, 50°05.07'N 114°36.47'W, sweeps, hillside seep, Carex utriculata , 17.vii.2011, 1 ♂, K. N. Barber leg. ( CNCI); Leduc, mixed grassland, 26.vi.1967, 1 ♀, D. Chomyn leg. ( PMAE); Nordegg, 25.vii.1936, 1 ♂, E. H. Strickland leg. ( USNM); Peace River, NE side of Peace R., sweep south facing grass slope, 13.vii.1997, 1 ♀, T. A. Wheeler leg. ( LEMQ 0039964); Peter Lougheed P. Pk., 50°41.82'N 115°06.92'W, sweeps, fen, Carex utriculata with cf. Poa pratensis , 13.vii.2011, 4 ♂♂ 4 ♀♀ ( DEBU 01502921–28), 14.vii.2011, 1 ♂ 1 ♀ ( DEBU 01503290, -91), 16.vii.2011, 3 ♂♂ ( DEBU 01503387–89); same locality but 50°41.85'N 115°06.93'W, sweeps, roadside grasses and herbs, 13.vii.2011, 3 ♂♂ 3 ♀♀ ( DEBU 01503281–86); same locality but Upper Lake Drive, 50°36.98'N 115°07.13'W, sweeps, edge of fen, Carex utriculata , 14.vii.2011, 1 ♀ ( DEBU 01503313), 16.vii.2011, 11 ♂♂ 5 ♀♀ ( DEBU 01503492–507), 21.vii.2011, 1 ♂ ( DEBU 01503671), sweeps/pooter, edge of fen, Carex utriculata , 15.vii.2011, 3 ♂♂ 2 ♀♀ ( DEBU 01503360–64); same locality but Upper Lake Drive, 50°36.94'N 115°07.16'W, sweeps, edge of fen, Carex utriculata , 16.vii.2011, 1 ♂ ( DEBU 01503646); same locality but 0.5 km W Little Highwood Pass, 50°38.55'N 115°02.94'W, sweeps, roadside grasses incl. Bromus inermis , 14.vii.2011, 1 ♂ 7 ♀♀ ( DEBU 01502558–65), 16.vii.2011, 27 ♂♂ 26 ♀♀ ( DEBU 01502699–751), all K. N. Barber leg.; Ponoka, 12.vi.1921, 1 ♂, O. Bryant leg. ( USNM); ~ 20 km SSE Robb, Hwy #40, Lovett River P[rovincial] R[ecreation] A[rea], 53°03.89'N 116°49.02'W, sweeps, streamside, mostly grasses, 23.vii.2008, 16 ♂♂ 8 ♀♀, K. N. Barber leg. ( DEBU); Rocky Mtn.House, 52°22.57'N 114°56.34'W, pooter, grasses on bridge embankment, edge of parking area, 24.vii.2008, 1 ♂; ~ 40.8 km SSW Rocky Mtn. House, Hwy #752, 52°09.79'N 115°25.89'W,sweeps/pooter, roadside Poeae , 24.vii.2008, 1 ♀, both K.N. Barber leg. (both CNCI); Sheep Creek P. Pk., 54°03.6'N 119°00.7'W, sweep at campground, 22.vii.2003, 2 ♂♂ ( LEMQ 0040455, -57); Kananaskis, Sheep River P. Pk., Sandy McNabb camp, 50°38.27'N 114°31.7'W, sweep, open forest & grasses, 28.vii.2003, 1 ♂ 1 ♀ ( LEMQ 0040397, -407), all S. Boucher leg.; Slave L., 14.viii.1924, 4 ♂♂, O. Bryant leg.

( CASC 2 ♂♂, USNM 2 ♂♂); same locality but 50°48.98'N 115°09.80'W, sweeps, graminoids and

herbs nr. edge of fen, 13.vii.2011, 1 ♂ 1♀ ( DEBU 01502859, -60); same locality but Buller Mt. day use, 50°52.02'N 115°21.23'W, sweeps, edge of fen, Carex utriculata ?, 15.vii.2011, 1 ♂ ( DEBU 01503379); W. A. Switzer P. Pk., Beaver Ranch Trail, 53°29.67'N 117°48.00'W, sweeps, sedge-nlled oxbow, Carex utriculata ?, 22.vii.2011, 1 ♂ ( DEBU 01503776); same locality but 53°29.80'N 117°48.02'W, sweeps, edge of Jarvis Ck., Carex utriculata , 22.vii.2011, 1 ♂ 2 ♀♀ ( DEBU 01503704–06), all K. N. Barber leg.; Wabamun, 1.vii.1966, 2 ♂♂, K. A. Spencer leg. ( BMNH, 1 ♂ genit. prep.); Waterton Lakes N. P., 7–12.vii.1980, 1 ♀, 14–20.vii.1980, 1 ♀, H. J. Teskey leg. ( CNCI); 10 km N Whitecourt, Sakwatamau R., 54°12'03"N 115°46'40"W, sweep sedges & grass at upper beach, 18.vii.2003, 1 ♀, T. A. Wheeler leg. ( LEMQ 0040424). BRITISH COLUMBIA: Adams Lake, 19.vi.1950, 1 ♀, F. Cameron leg. ( CNCI); Atlin, 2200', 3.vi.1955, 2 ♀♀, 4.vi.1955, 1 ♂ 1 ♀, 20.vi.1955, 1 ♂ 4 ♀♀, 21.vi.1955, 1 ♂ 1 ♀, 23.vi.1955, 1 ♂ 1 ♀, 24.vi.1955, 2 ♂♂ 1 ♀, 27.vi.1955, 3 ♀♀, 29.vi.1955, 1 ♀, 6.vii.1955, 1 ♀, 7.vii.1955, 1 ♂ (genit. prep.), 8.vii.1955, 1 ♀, B. A. Gibbard leg., 20.vi.1955, 1 ♂ 3 ♀♀, 23.vi.1955, 1 ♂ 3 ♀♀, 3.vii.1955, 1 ♀, 6.vii.1955, 1 ♂, H. Huckel leg. ( CNCI); S of Atlin, Warm Springs, sweep vegetation at warm stream margin, 2. vi.1997, 3 ♂♂ 3 ♀♀ ( LEMQ 0039914, -24, -25, -38, -39, 0040011); 23 km S Atlin, Warm Spring, 59°23'N 133°32'W, sweep vegetation stream, 25.vi.1997, 2 ♂♂ ( LEMQ 0040005, -06), all T. A. Wheeler leg.; Bear Lake, 20.vii.1903, 1 ♂ 2 ♀♀, R. P. Currie leg. ( USNM); Canim Lake, 22.vi.1938, 1 ♀, J. K. Jakob leg., 22.vi.1938, 2 ♀♀, G. S. Walley leg. ( CNCI); Chetwynd, 615 m, 25.vi.1978, 4 ♂♂ 1 ♀, P. H. Arnaud Jr. leg. ( CASC, 1 ♂ genit. prep.); Clinton, 14.vi.1938, 1 ♀, G. S. Walley leg. ( CNCI); nr. Cranbrook, sweeps, 16.vii.1980, 1 ♂, G. Gibson leg. ( DEBU); Fernie, Annex Pk., 49°30'43"N 115°04'07"W, 2.viii.2010, 3 ♂♂ 2 ♀♀, S. M. Paiero leg. ( DEBU 00334405, -08, -14, -16, -26); Fernie, Annex Pk., 49°30.72'N 115°04.13'W, sweeps, wet ditch, mostly Equisetum arvense & E. laevigatum , 17.vii.2011, 1 ♂ 3 ♀♀, sweeps, wet ditch, Carex utriculata , 17.vii.2011, 1 ♂ 1 ♀, K. N. Barber leg. ( CNCI); 35 km NW Fraser on Klondike Hwy, Tutshi Lake, sweep grasses beside road, 3.vi.1997, 2 ♀♀, T. A. Wheeler leg. ( LEMQ 0040014, -15); Glacier, 1.viii.1950, 1 ♀, K. Bourns leg. ( CNCI); Glacier National Park, ca. 1.6 km N Glacier, 14. vii.1974, 1 ♂ 2 ♀♀, P. H. Arnaud Jr. leg. ( CASC); Haines Cut-Off, 900', 13.vii.1948, 1 ♂ 1 ♀, W. R. Mason leg.; Honeymoon Bay, Cowichan L., 6.vi.1955, 1 ♀, G. E. Shewell leg.; 15 mi E Hope, 12.vii.1973, 2 ♀♀, H. J. Teskey leg. (all CNCI); Kaslo, 11.vi. [-], 2 ♀♀, H. G. Dyer leg., 11.vi. [-], 2 ♂♂ 2 ♀♀, 25.vi. [-], 1 ♀, R. P. Currie leg.; Kaslo Cr., 18.vi. [-], 2 ♂♂ 2 ♀♀, R. P. Currie leg., 18.vi. [-], 1 ♀, A. N. Caudell leg. (all USNM); Kimpton Creek, Hwy 93, 23.vi.1982, 1 ♂ 2 ♀♀, B. V. Peterson leg. ( CNCI); Kinbasket Lake, BC Hydro drawdown study, Malaise trap ( WIL 84-01), 18.vii.2008, 1 ♀, (33 MTRT 15), 12–13.vii.2009, 1 ♀, (34 MTRT 15), 12–13.vii.2009, 1 ♀, (84 MCOT 15), 21.vi.2010, 1 ♀, (87 MTRT 01), 29–30.vii.2009, 1 ♀, (88 MCOT 01), 10.vii.2010, 1 ♀, (88 MCOT 15), 10.vii.2010, 1 ♂, Cooper Beauchesne & Assoc. Ltd. leg. ( RBCM); ~ 7.9 km SE Valemount, edge of Kinbasket Lake, 52°46.70'N 119°10.44'W, sweeps, mixed graminoids & Equisetum , 23.vii.2011, 1 ♂, K. N.Barber leg.; King Salmon L., 58°43'N 132°54'W, 1750', Carex , grass, Equisetum , beside lake, 3.vii.1960, 1 ♂, W. W. Moss leg., around camp veg‘n, garbage, 14.vii.1960, 1 ♀, R. Pilfrey leg. (all CNCI); Kiskatinaw Public Campground, Alaska Hwy, DC-20 (31.4km), 26.vi.1978, 1 ♂ 1 ♀, P. H.Arnaud Jr. leg. ( CASC); Kootenay N. P., Daer-Pitts, Aspen Control 2, Malaise, 16–30.vii.2000, 1 ♂, G. Gareau leg. ( BDUC); Lac La Hache, 15.vii.1973, 1 ♂; Lac La Jeune, 27.vi.1973, 1 ♀, both H. J. Teskey leg.; Liard Hot Spring, mi. 496, 1500', Alaska Hwy, 9–10.vii.1959, 1 ♀, R. E. Leech leg.; Mission City, 6.vi.1953, 1 ♂, 18.vii.1953, 1 ♀, W. R. M. Mason leg., 27.vii.1953, 1 ♂, G. J. Spencer leg. (all CNCI); Mt. Robson

Prov. Pk., Hwy #16, small road towards Mt. Robson, 53°03'N 119°15'W, forest noor, swamp, (Universität Bielefeld, Ca1519), 6.viii.2002, 2 ♂♂; Mt. Robson Prov. Pk., hiking trail along Robson R. (53°02'06" N, 119°13'50" W) up to Kinney Lake (53°04'56" N, 119°11'05" W, 981 m), swept on herbs & shrubs incl. Carex spp., aspirated, (Universität Bielefeld, Ca1520), 6.viii.2002, 1 ♂; Nancy Greene Lake (Blueberry Creek), S of Hwy #3, ~ 24 km WSW Castlegar, ~ 49°14'N ~ 118°W, swept, aspirated, (Universität Bielefeld, Ca1534), 14.viii.2002, 1 ♂ 1 ♀, all M. v. Tschirnhaus leg. (all ZSMC, in ethanol); Nancy Greene Park, sweep, [-]. viii.1980, 1 ♂ 1 ♀, S. Marshall leg. ( DEBU); Prophet River Prov. Campground, Alaska Hwy, DC-221, 13.viii.1978, 1 ♂, P.H.Arnaud Jr. leg. ( CASC); Quesnel, 12.vi.1949, 1 ♀, G. J. Spencer leg. ( UBCZ, head glued to minuten); Revelstoke, 2.vii.1973, 2 ♀♀, H. J. Teskey leg.; Robson, 24.vi.1949, 1 ♀, 10.vi.1950, 1 ♀, 7.vii.1950, 2 ♂♂, H. R. Foxlee leg. (all CNCI); Robson, 28.v.1958, 1 ♀, H. R. Foxlee leg. ( UBCZ, genit. prep.); Salmon Arm, 17.vi.1925, 1 ♀, A. A. Dennys leg.; Sawmill Lk., Telegraph Ck., 1100', Carex , grass, Equisetum , beside lake, 2.vii.1960, 1 ♀, R. Pilfrey leg.; Summit Lake, mi. 392 Alaska Hwy, 4500', 8.vii.1959, 1 ♂ 1 ♀, E. E. MacDougall leg., 8.vii.1959, 1 ♂, 11–14.vii.1959, 1 ♀, R. E. Leech leg. (all CNCI); Alaska Hwy at Summit Lake, 58°39.1'N 124°39.1'W, sweep along small outlet stream, 12.vii.1997, 1 ♀, T. A. Wheeler leg. ( LEMQ 0039942); Alaska Hwy at Summit Lake Campground, sweep vegetation, 26.vii.2000, 1 ♂ 1 ♀, V. Dion & S. Boucher leg. ( LEMQ 0039915, -55); Terrace, 5.vi.1960, 1 ♂, C. H. Mann leg., 9.vii.1960, 1 ♀, 21.vii.1960, 1 ♀, 24.vii.1960, 1 ♀, W. R. Richards leg.; Gagnon Rd., 6 mi W Terrace, 220', on buttercup, 20.vi.1960, 1 ♂ 2 ♀♀; same locality but on blueberry, 20.vi.1960, 1 ♀, all W. W. Moss leg.; Kitsumkalum, Lake, 20 mi N Terrace, 500', 16.vi.1960, 1 ♀, R. Pilfrey leg.; Kleanza Ck., 14 mi E Terrace, 14.vii.1960, 4 ♂♂ 3 ♀♀, G. E. Shewell leg.; same locality but 250', 17.vii.1960, 1 ♂, R. Pilfrey leg.; Remo, 7 mi E Terrace, 230', 13.vi.1960, 2 ♀♀, R. Pilfrey leg.; Shames, 17 mi W Terrace, 13.vii.1960, 1 ♂, J. G. Chillcott leg.; Shames, 18 mi SW Terrace, 23.vi.1960, 1 ♀, C. H. Mann leg.; Zymagotitz River, 6 mi W Terrace, 190', 20.vi.1960, 1 ♂, R. Pilfrey leg.; Tunjony L., 58°26'N 132°45'W, 3200', 20.vii.1960, 1 ♀, W. W. Moss leg. (all CNCI); Vancouver Is., Courtenay, [-]. vi.1965, 1 ♂, N. L. H. Krauss leg. ( USNM); Williams L., 20.viii.1960, 4 ♂♂ 1 ♀, W. R. Richards leg.; 20 km SW Williams Lake, CD1420, 20.vii.1992, 3 ♂♂ 2 ♀♀ (incl. pair in copula), A. Borkent leg. (all CNCI); Yarrow, NW foot of Vedder Mt., 49°04'N 122°03'W, swept/aspirated, herbs along forest road, (Universität Bielefeld, Ca1509), 23.vii.2002, 1 ♀, M. v. Tschirnhaus leg. ( ZSMC, in ethanol). LABRADOR: Cartwright, 26.vi.1955, 1 ♀, E. E. Sterns leg.; Hopedale, 8.viii.1926, 1 ♀, W. W. Perrett leg. (both CNCI) . MANITOBA: Aweme, 4.vii.1917, 1 ♂, N. Criddle leg., 16.vii.1923, 1 ♂, H. A. Robertson leg. ( CNCI); Aweme, Criddle farm, 49°42.5'N 99°36.1'W, sweep open forest at graveyard, 8.vii.2000, 1 ♂ 1 ♀, T. A. Wheeler leg. ( LEMQ 0039922, -30); Churchill, 12.vii.1952, ecological data T-C 3-1, 1 ♂, J. G. Chillcott leg. ( CNCI); 11 km S Churchill, Goose Creek Bridge, 58.662°N 94.165°W, 0 m a.s.l., 16.vii.2010, 1 ♂, J. Wang leg. ( BIOUG, in ethanol, Barcode of Life, Sample ID:10PROBE-14693, BOLD ID: JWDCJ 353-11); 26 km SE Churchill,Twin Lakes, 58.63°N 93.819°W, 14 m a.s.l., 19.vii.2009, 1 ♀, Arctic Ecology 2009 leg.( BIOUG, in ethanol, Barcode of Life, Sample ID:09PROBE-JW0720, BOLD ID: JWDCC 720-10); 26 km SE Churchill, Twin Lakes, burn site, 58.6173°N 93.8123°W, 38 m a.s.l., 1.viii.2010, 1 ♂, J. Wang leg. ( BIOUG, in ethanol, Barcode of Life, Sample ID:10PROBE-12875, BOLD ID: JWDCH 669-10); Churchill, 12 km E on Launch Rd., 58°45'16"N 93°59'55"W, fen, Malaise trap, 10–14.vii.2007, 1 ♂, A. Renaud leg. ( DEBU 00362140); 13 km E Erickson, 31. vii.1983, 1 ♂, K. N. Barber leg. ( DEBU); Ninette, maple / elm noodplain community, along stream, 25.vii.1958, 1 ♂, N. B. Chillcott leg.; 5 mi SW Shilo, 28.v.1958, 1 ♀, C. D. F. Miller leg.; 5 mi SW Shilo, noodplain community near tamarack bog, 28.v.1958, 2 ♀♀; 5 mi SW Shilo, noodplain community near tamarack bog, sedge meadows, 16.vi.1958, 1 ♀; same locality but ex. Anemone sp., 6.viii.1958, 1 ♀, all J. F. McAlpine leg.; same locality but swampy wooden hillside, 2.viii.1958, 1 ♂, J. G. Chillcott leg. (all CNCI). NEW BRUNSWICK: Chamcook, 30.vi.1965, 1 ♀; Chamcook, Glebe Road, 28.vi.1965, 1 ♂, both G. E. Shewell leg. (both CNCI). NOVA SCOTIA: C[ape] B[reton] H[ighlands] N. P., Mackenzie Mt., PG640848, dry spruce birch forest, 24.vi.1984, 2 ♀♀, H. J. Teskey leg. ( CNCI); Cape Breton Island, 1 km N Mabou, sweep grasses in roadside picnic area, 16.viii.1996, 1 ♀, T. A. Wheeler leg. ( LEMQ 0039928). NORTHWEST TERRITORIES: Fort Liard, 9.vi.1969, 1 ♀, G. E. Shewell leg.; Ft. McPherson, 2. vii.1957, 1 ♂, S. D. Hicks leg., 10.vii.1957, 1 ♂, R. Hurley leg. (all CNCI); Fort Smith, 60°00.109'N 111°51.508'W, frog swamp, sweep, 29.vi.2012, 2 ♂♂, S. & K. Williamson leg. ( BDUC); Norman Wells, 2.vii.1969, 1 ♂, G. E. Shewell leg. ( CNCI); Norman Wells, 65.26694°N 126.72827°W, 62 m, Repl. 2 mesic, sweep, 17.vi.2011, 1 ♂ 1 ♀, NBP Field Party leg. ( LEMQ); Norman Wells, nr. D.O.T. Lake, 65.252°N 126.665°W, sweep, 12.vi.2011, 2 ♂♂ 1 ♀, A. M. Solecki leg. ( LEMQ). ONTARIO: Algonquin Pk., 4.vii.1978, 1 ♀, D. McCorquodale leg.; Algonquin P. Pk., nr. Madawaska Lake, 45.3261°N 78.3056°W, Malaise trap, hardwood forest ( MD-GAP), 21.vi.–4.vii.2008, 1 ♀, E. Proctor leg. (both DEBU); Algonquine [sic], mixed wood, 1.vi.1991, 1 ♂ 2 ♀♀ (1 ♀ genit. prep.); Beaver Bay, mixed wood nr. river, 31.v.1991, 1 ♂ (genit. prep.), all M. Barták leg. (all MBPC); Bruce Co., Crane River & Hwy #6, sweep, 23.vi.1995, 1 ♀, S.A. Marshall leg. ( DEBU); Bruce Peninsula N. P., Bartley Lake, 45°12.9'N 81°29.3'W, sweeps, shoreline graminoids, 2.vii.1999, 1 ♀; Bruce Peninsula N. P., Cameron Lk. Rd., 45°12.5'N 81°33.5'W, sweeps, trailside grasses, mixed forest, 4.vii.1998, 6 ♂♂ 1 ♀, sweeps, trailside veg., grasses, mixed forest, 4.vii.1998, 2 ♂♂ 2 ♀♀; same locality but 45°12.7'N 81°33.0'W, sweeps, roadside vegetation, 5.vii.1998, 1 ♀; Bruce Peninsula N. P., Emmett Lake, 45°13.5'N 81°28.2'W, sweeps, mostly graminoids, open area under Acer / Quercus , 2.vii.1999, 2 ♂♂ 6 ♀♀, all K. N. Barber leg. (all DEBU); Charlton, 29.vii.1930, 1 ♀, H. S. Parish leg. ( USNM); Cochrane, 49°03.54'N 81°04.41'W, sweeps, grasses on edge of hwy.pulloff, 21.vii.2009, 5 ♂♂ 2 ♀♀, K. N. Barber leg. ( CNCI); Dubreuilville, 48°21.09'N 84°33.90'W, sweeping undergrowth Pinus / Populus forest, with Clintonia , Vaccinium , ferns, graminoids, 10.vii.2010, 4 ♂♂, J. Roháček leg. ( SMOC, 1 ♂ genit. prep.); Dubreuilville, 48°21.12'N 84°34.04'W, sweeps, Equisetum ssuviatile , Carex , along Magpie River, 10.vii.2010, 1 ♂; ~ 35 km WSW Dubreuilville, 2 km SE jct. Hwys #17 & #519, 48°17.16'N 84°53.34'W, sweeps, roadside vegetation incl. wet ditch, 31.vii.2008, 1 ♀, both K. N. Barber leg.; Eagle River, 11.vii.1960, 1 ♀, Kelton & Whitney leg.; ~ 55 km NNW Elliot Lake, s. of Rocky Island Lake, 46°49.32'N 82°59.54'W, sweeps, Scirpus , grasses, Equisetum , ferns, 3.vii.2010, 3 ♂♂ 2 ♀♀, K. N. Barber leg. (all CNCI); same locality but 455 m, sweeping vegetation with predominant grasses, 3.vii.2010, 11 ♂♂ 17 ♀♀ (1 ♂ 1 ♀ genit. prep.); ~ 61 km NNW Elliot Lake, Three Lakes, 46°49.94'N 83°06.31'W, 425 m, sweeping grasses [mostly Glyceria striata ] near lakeshore, 3.vii.2010, 7 ♂♂ 6 ♀♀ (1 ♀ genit. prep.), all J. Roháček leg. (all SMOC); ~ 63 km NNW Elliot Lake, s. of Rocky Island Lake, 46°49.80'N 83°09.08'W, sweeps, streamside graminoids, 4.vii.2010, 3 ♂♂ 3 ♀♀, K. N. Barber leg. ( CNCI); same locality but 425 m, sweeping streamside graminoids, 4.vii.2010, 4 ♂♂ 2 ♀♀, J. Roháček leg. ( SMOC); ~ 66 km NNW Elliot Lake, Rocky Island Lake, 46°50.82'N 83°08.76'W, sweeps, Scirpus [ microcarpus ] on dried shoreline, 4.vii.2010, 1 ♂, K. N. Barber leg.; Finland, 21.vii.1960, 1 ♂, S. M. Clark leg. (both CNCI); Fork[s of the] Credit, 2.viii.1933, 1 ♀, H. S. Parish leg. ( USNM); Foymont, 1600', 25.vi.1983, 1 ♀, H. Walther leg. ( CNCI); Greenwater P. Pk., 49°10.93'N 81°16.37'W, sweeps, Phalaris arundinacea in creek noodplain, 21.vii.2009, 1 ♂ ( DEBU 01502119); Greenwater P. Pk., Sandbar Lk.Trail, 49°13.10'N 81°17.35'W, sweeps, lakeshore Equisetum spp., graminoids, Caltha , 21.vii.2009, 1 ♀ ( DEBU 01502089), both K. N. Barber leg.; ~ 5 km NE Heyden, Hwy 552, 46°41.37'N 84°16.85'W, sweeps, roadside ditch/ stream, graminoids, 31.vii.2006, 1 ♀; 12 km NE Heyden, Bellevue Tower Hill, open meadow & mixed forest, 4. vi.1987, 1 ♀, both K. N. Barber leg.; Icewater Creek WS [watershed], ~12.7 km NNE Searchmont, mi.10.5 Whitman Dam Rd., riparian meadow/ alder thicket, Malaise, 29.vi.–3.vii.1986, 1 ♀, 10–15.vii.1986, 2 ♂♂, 15–21.vii.1986, 1 ♀, [K. N. Barber] leg.; same locality but 13.5 km NNE Searchmont, mi.11.5 Whitman Dam Rd., sandy access road, 20.vi.1986, 6 ♂♂ 6 ♀♀, 1.vii.1986, 3 ♀♀, 15.vii.1986, 1 ♀; Icewater Creek WS [watershed], 46°53.7'N 84°03.4'W, sweeps, roadside sedges / grasses, mixed forest, 10.vii.1998, 3 ♂♂ 3 ♀♀, sweeps, roadside sedges / grasses, mixed forest, 7.vii.1998, 4 ♂♂ 4 ♀♀, sweeps, Thalictrum , Eupatorium [ Eutrochium ], sedges in mixed forest, 17.vii.1998, 1 ♂, sweeps, Thalictrum , sedge, fern, riparian mixed forest, 7.vii.1998, 2 ♂♂ 2 ♀♀, sweeps, trailside sedges, ferns, grasses, 10.vii.1998, 2 ♂♂, sweeps, trailside vegetation incl. sedges, ferns, grasses, 24.vii.1998, 1 ♀; same locality but 46°53.72'N 84°03.39'W, sweeps, riparian ferns, graminoids, 5.viii.2006, 8 ♂♂ 2 ♀♀, all K. N. Barber leg.; Iroquois Falls, Carex , grass, on unshaded wet soil, 24.vi.1987, 1 ♂, swept along small cold mossy stream, 29.vi.1987, 3 ♂♂, J. R. Vockeroth leg. (all CNCI); Kirkland Lake, 22.vi.1980, 1 ♂, 24.vi.1980, 1 ♀, J. Cashaback leg. ( DEBU); Lake Superior P. Pk., 47°16.42'N 84°33.66'W, pooter, grasses in roadside ditch, 15.vi.2004, 2 ♂♂ 1 ♀, K. N. Barber leg. ( DEBU 01501450–52); Lake Superior Prov. Park, Crescent Lake Trail, 47°16.81'N 84°33.23'W, sweeping, mostly graminoids under Betula / Acer , 9.vii.2010, 8 ♂♂ 5 ♀♀, J. Roháček leg. ( SMOC, 1 ♂ 1 ♀ genit. prep., 1 ♂ 2 ♀♀ photographed); Lake Superior P. Pk., Trappers Trail, 47°32.53'N 84°48.50'W, sweeps, trailside sedges, 13.vi.2010, 5 ♂♂ 11 ♀♀, K.N. Barber leg.( DEBU 01502469–84); Manitoulin Is., Carter Bay, 45°36.3'N 82°08.5'W, sweeps, creekside graminoids, 30.vi.1999, 2 ♂♂, sweeps, Pearly Everlasting [ Anaphalis margaritacea ], 30.vi.1999, 1 ♂ ( CNCI); Manitoulin Is., 0.7 km N Michael’s Bay Pk., 45°36.4'N 82°06.1'W, sweeps, low veg. in mixed wood, 5.vii.1998, 16 ♂♂ 12 ♀♀ ( CNCI 12 ♂♂ 8 ♀♀, 1 ♂ genit.prep., NMPC 2 ♂♂ 2 ♀♀, SMOC 2 ♂♂ 2 ♀♀), 4.vii.1999, 4 ♂♂ 10 ♀♀ ( CNCI, 1 ♀ genit. prep.), all K. N. Barber leg.; Manitoulin Is., Poplar, 45°46'N 82°28'W, sweep along grassy trail, 27.vi.1992, 1 ♀, T. A. Wheeler leg. ( LEMQ 0039917); Hwy #17, ~ 8.5 km NW Marathon, 48°47.69'N 86°26.07'W, sweeps, roadside graminoids, 7 ♂♂ 5 ♀♀, 31.vii.2008, K. N. Barber leg.; Maynooth, 18.vi.1963, 1 ♂, J. F. McAlpine leg.; Minnitaki, 13.vi.1960, 1 ♀, Kelton & Whitney leg. (all CNCI); Moosonee, 51.24690°N 80.68102°W, Repl. 3 mesic, pan traps, 17–20.vi.2010, 1 ♀, 20–23.vi.2010, 1 ♀; Moosonee, 51.28288°N 80.63926°W, Repl. 2 wet, pan traps, 19–23.vi.2010, 1 ♂, all NBP Field Party leg. (all LEMQ); Moosonee, 51°14.79'N 80°40.35'W, sweeps, mostly Solidago , Calamagrostis , 9.vii.2014, 1 ♀; Moosonee, 51°16.33'N 80°39.11'W, sweeps, mostly Rubus , Impatiens , under Salix , Alnus , 10.vii.2014, 2 ♂♂ 1 ♀, 11.vii.2014, 4 ♂♂ 1 ♀; Moosonee, 51°16.36'N 80°39.11'W, sweeps, railside ditch, mostly Equisetum ssuviatile , Carex spp., 10.vii.2014, 4 ♀♀, 11.vii.2014, 1 ♀; Moosonee, 51°16.54'N 80°39.00'W, sweeps, Equisetum , Rubus , Cornus , graminoids, edge of wet forest trail, 8. vii.2014, 1 ♂ 1 ♀, 9.vii.2014, 1 ♂ 6 ♀♀, 10.vii.2014, 8 ♂♂ 20 ♀♀, 11.vii.2014, 7 ♂♂ 13 ♀♀; Moosonee, 51°16.55'N 80°39.01'W,sweeps, mostly Carex spp., wet forest trail, 11.vii.2014, 4 ♂♂ 3 ♀♀; Moosonee, 51°16.63'N 80°38.87'W, sweeps, Calamagrostis , Carex , drier edge of sedge meadow, 9.vii.2014, 1 ♀; Moosonee, 51°16.68'N 80°38.65'W, sweeps, mostly C [arex] utriculata , C. aquatilis , wet sedge meadow, 10.vii.2014, 1 ♀; Moosonee, 51°16.75'N 80°38.76'W, sweeps, Equisetum & graminoids, wet edge of sedge meadow, 9.vii.2014, 1 ♂; Moosonee, 51°16.99'N 80°38.37'W, sweeps, mostly Equisetum ssuviatile , Carex spp., wet sedge meadow, 10.vii.2014, 1 ♂ 2 ♀♀, all K. N. Barber leg. (all CNCI); Nestor Falls, 7–8.viii.1960, 1 ♀, Kelton & Whitney leg.; ~ 8 km SSW Nipigon, Hwy #628, 48°57.09'N 88°19.71'W, sweeps, damp roadside, mixed graminoids, 16.vii.2008, 1 ♀; ~ 20 km E Nipigon, Hwy #17, rest area, 48°58.00'N 87°59.09'W, sweeps, Aster [ Eurybia ], Rubus , Aralia , Diervilla , 31.vii.2008, 2 ♂♂ 3 ♀♀; Otter Rapids, 50°11.08'N 81°38.37'W, sweeps, Eurybia , Equisetum under Populus , 19.vii.2009, 7 ♂♂ 8 ♀♀, 20.vii.2009, 8 ♂♂ 6 ♀♀; Otter Rapids, 50°10.91'N 81°38.54'W, sweeps, mostly Calamagrostis canadensis in hydro cut, 19. vii.2009, 1 ♂ 4 ♀♀, sweeps, mostly C. canadensis , E. ssuviatile in hydro cut, 20.vii.2009, 5 ♂♂ 6 ♀♀; Otter Rapids, 50°10.96'N 81°37.88'W, sweeps, grasses, herbs on roadside slope, 20.vii.2009, 1 ♀, all K. N. Barber leg. (all CNCI); Pancake Bay P. Pk., 46°57.74'N 84°42.63'W, sweeps, beach grasses [ Ammophila breviligulata ], 7.viii.2004, 6 ♂♂ 2 ♀♀ ( DEBU 01501206–13); Pancake Bay P. Pk., 46°58.11'N 84°42.62'W, sweeps, open grassy area, 17.vii.2004, 1 ♂ ( DEBU 01500263); Pancake Bay P. Pk., 46°58.13'N 84°42.70'W, sweeps, open grassy area near highway, 17.vii.2004, 1 ♂, 7.viii.2004, 4 ♂♂ 1 ♀ ( DEBU 01501103–07), all K. N. Barber leg.; Hwy #17 at Prairie River mouth, 48°48.32'N 86°46.64'W, sweeps, grasses, composites, Rubus , forest edge, 12.viii.2006, 3 ♂♂, 16.vi.2007, 4 ♂♂ 3 ♀♀, K. N. Barber leg. ( CNCI); Pukaskwa N. P., Coastal Trail, Hattie Cove–Playter Harbour, sweep, 21. vii.2001, 1 ♀, M. Buck leg. ( DEBU 00183046); René Brunelle P. Pk., 49°25.89'N 82°08.44'W, sweeps, roadside Equisetum spp., 19.vii.2009, 1 ♂ 1 ♀, K. N. Barber leg. ( DEBU 01502000–01); ~ 2 km E Rossport, Hwy #17 picnic area, 48°50.3'N 87°29.4'W, 9.vii.1999, sweeps of graminoids, 1 ♂ 2 ♀♀; S[ault] S[ainte] Marie, S. of Algoma U[niversity] College, 46°29.9'N 84°17.2'W, sweeps, graminoids under Populus / Betula , 11.vii.1997, 1 ♀, sweeps, grassy edge of Populus / Betula , 11.vii.1997, 1 ♀, sweeps, grassy edge of woods, 11.vii.1997, 1 ♀, sweeps, trailside Carex / Scirpus , 14.vii.1998, 3 ♂♂ 2 ♀♀, all K. N. Barber leg. (all CNCI); S[ault] S[ainte] Marie, Baseline Rd., 46°31.40'N 84°24.40'W, Malaise #1, Aster [ Doellingeria ], Rubus , Equisetum , Carex , Solidago , in aspen clearing, 7–17.vi.2005, 1 ♀, 17–29.vi.2005, 1 ♂, 18–29.vii.2005, 1 ♂, sweeps, Rubus , Equisetum , Thalictrum , Impatiens , ferns, under aspen, 17.vi.2005, 1 ♂, sweeps, Aster [ Doellingeria ], Rubus , Equisetum , Carex , ferns, under aspen, 22.vi.2005, 1 ♂ 1 ♀, 25.vi.2005, 1 ♂, 26.vi.2005, 1 ♂ 2 ♀♀, 14.vii.2005, 2 ♂♂ 1 ♀, 18.vii.2005, 1 ♂ 1 ♀, 19. vi.2011, 1 ♀, sweeps, mostly ferns under aspen, 19.vi.2011, 1 ♀, sweeps, mostly Carex in aspen clearing, 25.vi.2005, 1 ♂ 1 ♀, K. N. Barber leg. ( CNCI); same locality but 46°31.40'N 84°24.45'W, sweeps, edge of forest, Solidago , Rubus , Equisetum , grasses, 25.vi.2005, 2 ♂♂ 7 ♀♀, 26.vi.2005, 1 ♂ 6 ♀♀ ( CNCI), 10.vii.2005, 9 ♂♂ 5 ♀♀ ( CNCI 7 ♂♂ 3 ♀♀, 1 ♂ wing illustration, SMOC 2 ♂♂ 2 ♀♀), 14.vii.2005, 5 ♂♂ 3 ♀♀, 29.vii.2005, 2 ♀♀ ( AMNH), K. N. Barber leg.; same locality but w. of creek, 46°31.52'N 84°24.63'W, sweeps, Carex under ash / aspen, 22.vii.2005, 1 ♀, sweeps, graminoids on path, 22.vii.2005, 1 ♀; same locality but w. of creek, 46°31.61'N 84°24.68'W, sweeps, graminoids/ composites, 20.viii.2008, 1 ♂, both K. N. Barber leg. (both CNCI); S[ault] S[ainte] Marie, Birchwood Pk., mixed forest, 28.vi.1986, 1 ♂, 6.vii.1986, 3 ♀♀, 26.vii.1986, 1 ♀; same locality but 46°30.7'N 84°15.6'W, sweeps, including Impatiens under Betula / Acer , 20.vi.1998, 1 ♂, sweeps, Impatiens under Betula / Acer , 27.vi.1998, 1 ♂, sweeps, mostly sedges under Betula / Acer , 23.vi.1998, 1 ♂; S[ault] S[ainte] Marie, Bristol Place, 46°30.8'N 84°16.4'W, at MV [mercury vapour] lights in yard, 10.vii.2000, 1 ♀; S[ault] S[ainte] Marie, Bristol Pl. Pk., 46°30.8'N 84°16.6'W, [sweeps], graminoids, Impatiens , Rubus , Clematis , 18.vii.2006, 1 ♀, all K. N. Barber leg. (all CNCI); Sault Ste. Marie, Finn Hill, 46°31.48'N 84°17.36'W, sweeping graminoid vegetation, 7.vii.2010, 1 ♂ 1 ♀, J. Roháček leg. ( SMOC, 1 ♀ genit. prep.); S[ault] S[ainte] Marie, Finn Hill, 46°31.48'N 84°17.39'W, sweeps, graminoids, Equisetum , Rubus , Eurybia , 25.vii.2010, 2 ♂♂ 1 ♀; same locality but 46°31.49'N 84°17.39'W, sweeps, mostly grasses, 25.vii.2010, 1 ♂ 3 ♀♀, K. N. Barber leg. ( CNCI); Sault Ste. Marie, Fish Hatchery Road, near Coldwater Creek, 46°34.29'N 84°17.21'W, sweeping graminoids, Impatiens , 9.vii.2010, 1 ♀, J. Roháček leg. ( SMOC); S[ault] S[ainte] Marie,Ft.Creek Cons[ervation] Area, 46°32.5'N 84°20.8'W, sweeps, Carex under Populus / Acer , 30.vi.2002, 1 ♂, sweeps, trailside vegetation, mostly Poa sp., 8.vii.1998, 4 ♂♂ 4 ♀♀, sweeps, Impatiens , sedge, fern, 8.vii.1998, 1 ♀; S[ault] S[ainte] Marie, Kinsmen Pk., 46°35.5'N 84°16.6'W, sweeps, trailside graminoids in mixed forest, 7.viii.1997, 1 ♂ 2 ♀♀, all K. N. Barber leg. (all CNCI); S[ault] S[ainte] Marie, Sault Coll[ege] Outdoor Lab, 46°32.1'N 84°18.2'W, sweeps, low vegetation under Acer / Betula , 18.vi.1998, 2 ♂♂ 1 ♀, sweeps, trailside graminoids, Impatiens under Acer / Betula , 11.vi.1999, 1 ♂ 1 ♀, sweeps, short trail grasses, Acer / Betula , 25.vii.1997, 5 ♂♂ 6 ♀♀, sweeps, trailside grasses, Acer / Betula , 25.vii.1997, 2 ♂♂ 3 ♀♀, sweeps, trailside Carex under Acer / Betula , 26.vi.2002, 3 ♀♀, 29.vi.2002, 1 ♀, 30.vi.2002, 1 ♂ 1 ♀, sweeps, Carex sp., 25.vii.1997, 2 ♂♂; S[ault] S[ainte] Marie, hydro cut nr. Sault Coll[ege] Outdoor Lab, 46°32.1'N 84°18.0'W, sweeps, mostly sedges, 10.vii.2002, 1 ♂ 2 ♀♀, sweeps, Scirpus cyperinus , 12.viii.2008, 1 ♂; same locality but 46°32.18'N 84°18.01'W, sweeps, mostly Calamagrostis , Scirpus cyperinus , on trail, 12.viii.2008, 1 ♂, all K. N. Barber leg. (all CNCI); S[ault] S[ainte] Marie, Voyageur Trail, 46°35.48'N 84°15.23'W, sweeps, Scirpus cyperinus , 9.vii.2006, 1 ♀, [K. N. Barber] leg.; S[ault] S[ainte] Marie, Wishart Park, 46°33.86'N 84°17.57'W, sweeps, graminoids/ferns under canopy, 23.vii.2005, 1 ♀; 6 km ENE Searchmont,Whitman Dam Falls, sweeps, 1.vii.1986, 1 ♀, both K. N. Barber leg. (all CNCI); 12.4 km NNE Searchmont, mi.10 Whitman Dam Rd., herb/grass meadow by Goulais R., 1.vii.1986, 1 ♂ 2 ♀♀, K. N. Barber leg., 2. vii.1986, 1 ♀, 3.vii.1986, 1 ♂, 9.vii.1986, 1 ♂ 1 ♀, D. J. M. Harvey leg.; 18 km NNE Searchmont, mi.15 Whitman Dam Rd., grassy access road, 2.vi.1986, 1 ♂, 6.vi.1986, 1 ♀, D. J. M. Harvey leg., 6.vi.1986, 4 ♀♀, 19.vi.1986, 2 ♂♂, 24.vi.1986, 1 ♀, K. N. Barber leg.; same locality but Vaccinium / Sphagnum , 19.vi.1986, 1 ♀; ~ 18 km NNE Searchmont, Goulais River WS [watershed], ~mi. 15 Whitman Dam Rd., 46°55.3'N 83°56.3'W, sweeps, grass under jackpine, 30.vii.1999, 2 ♂♂ 5 ♀♀; ~ 21 km NNE Smooth Rock Falls, 49°20.91'N 81°32.01'W, sweeps, ditchside Equisetum spp., grasses, herbs, 19.vii.2009, 4 ♀♀, all K. N. Barber leg. (all CNCI); The Shoals P. Pk., Prairie Bee, 47°52.35'N 83°53.56'W, sweeps, grasses nr. parking area, 17.vi.2004, 1 ♂, K. N. Barber leg. ( DEBU 01500176); Algoma District,Thessalon, 18.vi.1965, 1 ♂, K.P. Butler leg. ( LEMQ 0039929); ~ 74 km NNE Thessalon, 46°53.94'N 83°16.23'W, shore of Mississagi R., sweeps, graminoids, herbs, Equisetum spp., 17.vii.2010, 1 ♂, K. N. Barber leg. ( CNCI); ~ 92 km NNE Thessalon, Mountain Ash Lake, 47°03.53'N 83°12.56'W, sweeping Clintonia , Eurybia , ferns, Diervilla under Betula , 5.vii.2010, 2 ♀♀, J. Roháček leg. ( SMOC); Thunder Bay, 22.vii.1970, 1 ♂, K. J. G. Deacon leg. ( DEBU, head glued to point); Timagami, 19.viii.1947, 1 ♀, G. S. Walley leg. ( CNCI); Waubamick [sic Waubamik], 14.vi.1915, 1 ♀, H. S. Parish leg. ( USNM); White River, 48°35.5'N 85°16.6'W, sweeps, mixed grasses, edge of parking lot, 9.vii.1999, 1 ♂ 3 ♀♀; White River, behind Continental Motel, 48°35.65´N 85°16.82'W, pooter, grasses, 16.vi.2004, 2 ♀♀; White River, “Black’s Cabin”, 48°36.71'N 85°15.61'W, sweeps, mowed grass/herbs on stable sand, 14.vi.2003, 1 ♀; 40 km SSW White River, 48°14.13'N 85°22.01'W, B6S4, boreal mixedwood, Malaise trap, 7–22.vii.2003, 1 ♂; same locality but 48°13.95'N 85°22.20'W, B6S1, boreal mixedwood, sweeps, Calamagrostis canadensis , 23.vii.2003, 1 ♂ 3 ♀♀, all K. N. Barber leg. (all CNCI). QUEBEC: Bolton Pass, Knowlton, 800', 5.vi.1963, 1 ♂ 1 ♀, J. R.Vockeroth leg.; Bradore Bay, 19.vii.1929, 1 ♀, W. J. Brown leg.; Breckenridge, 11.vi.1959, 1 ♀, C. H. Mann leg.; Fort Chimo [Kuujjuaq], 2.vii.1954, 3 ♀♀, W. R. Richards leg.; Gaspé, 22.vi.1954, 1 ♂, G. P. Holland leg. (all CNCI); Gaspé, ZEC [Zone d’Exploitation Contrôlée] York-Baillargeon, 48°49'N 64°52'W, shore of Lac Ross, sweep, 26.vii.2000, 1 ♂ ( LEMQ 0039927), forest edge near Lac Pauline, 26.vii.2000, 1 ♀ ( LEMQ 0039916), H. Varady-Szabo leg.; Gaspesie, l’Anse-au-Griffon, “La vallée” trail, sweep near marsh, 48°55'45"N 64°19'19"W, 14.viii.2006, 1 ♂ ( LEMQ 0040304); Gaspésie, near Routhierville, 48°10.9'N 67°08.8'W, sweep picnic area, 4.viii.2001, 1 ♀ ( LEMQ 0039916), both S. Boucher leg.; Great Whale River, 26.vi.1949, 1 ♀, 8.vii.1949, 1 ♀, 13.vii.1949, 1 ♀, J. R. Vockeroth leg. ( CNCI); Ile Bonaventure, 48°30'N 64°10'W, 3 km from Côte de Percé, sweep grass, 28.vii.2000, 1 ♂, H. Varady-Szabo leg. ( LEMQ 0039910); Indian House Lk., 9.vii.1954, 1 ♀, 18.vii.1954, 2 ♂♂, 27.vii.1954, 2 ♀♀, W. R. Richards leg.; La Verendrye P. Pk., mi.139 Rte.58, 28.vi.1965, 2 ♀♀, D. M. Wood leg. (all CNCI); Lac Roddic, 16 km S Maniwaki, 22.vi.1991, 4 ♂♂ 2 ♀♀, M. Barták leg. ( MBPC, 1 ♀ genit. prep.); Laniel, 17–18.vi.1963, 1 ♂, W. Gagne leg. ( CNCI); Laurentide Pk., 7.viii.1956, 1 ♀, A. H. Sturtevant leg. ( USNM); Magog, 1.vi.1965, 1 ♀, D. M. Wood leg.; Mistassini, 3.vii.1956, 1 ♀, 7.vii.1956, 1 ♂, J. R. McGillis leg.; Mistassini Post, 5.vii.1956, 1 ♀, 14.vii.1956, 1 ♂, 21.vii.1956, 2 ♀♀, J. R. Lonsway leg.; Mt. Albert, 11.viii.1954, 1 ♂, J. E. H. Martin leg.; Mt. Ste. Marie, Low, 1800', 22.vi.1965, 1 ♀, J. R. Vockeroth leg.; Newport, 14.viii.1983, 1 ♀, B. M. Bissett leg.; Old Chelsea, 28.v.1963, 1 ♀, 4.vi.1964, 1 ♀, J. R. Vockeroth leg.; Old Chelsea, King Mt., 26.v.1963, 1 ♀, J. G. Chillcott leg. (all CNCI); Old Chelsea, Summit King Mt., 13.vi.1980, 1 ♂, K. N. Barber leg. ( DEBU); Old Chelsea, Summit King Mt., 1150', 11.vi.1959, 3 ♂♂, 18.vii.1961, 1 ♂, 25.vi.1962, 3 ♂♂ 3 ♀♀, 2.vi.1963, 1 ♂ 1 ♀, 18.vi.1963, 1 ♂, 21.vi.1963, 1 ♂, 4.vi.1964, 1 ♀, 8.vi.1964, 2 ♂♂ 1 ♀, 15.vi.1964, 2 ♂♂ 4 ♀♀, 24.vi.1964, 1 ♂, 16.vi.1971, 6 ♀♀, J. R. Vockeroth leg. ( CNCI); Parc Jacques Cartier, Lac a la Chutes, 47°20'N 71°21'W, sweep open meadow, 2.viii.1999, 1 ♂, T. A. Wheeler leg. ( LEMQ 0040363); Kam[ouraska] Co., Parke Reserve, 950', 5.viii.1957, 1 ♂, E. Mason leg. ( CNCI). SASKATCHEWAN: Beaver Creek Cons[ervation] Area, ~ 13 km S Saskatoon, 51°58.6'N 106°43.0'W, sweeps, mostly grasses under Betula / Populus , 12.vii.1999, 10 ♂♂ 9 ♀♀, 17.vii.1999, 2 ♂♂ 1 ♀, K. N. Barber leg. ( CNCI); Beaver River, 5 mi S Goodsoil, 6.vii.1985, 1 ♂, D. W. Webb leg. ( INHS 40,219); Christopher Lake, [-]. v.1959, 1 ♂, A. & J. Brooks leg. ( CNCI); E side of Cowan Lake, 54°00'16"N 107°14'19"W, sweep near lake margin, 22.vii.2003, 1 ♂, T. A. Wheeler leg. ( LEMQ 0040410); Kenosee, 7.vi.1958, 2 ♀♀, A. R. Brooks leg.; ~6.5 km SE Langham, Hwy #16, 52°19.13'N 106°53.03'W, sweeps, mostly Elymus glaucus , 21.vii.2008, 3 ♂♂ 1 ♀, K. N. Barber leg.; Saskatoon, 19.vi.1926, 1 ♀, 29.vi. 192[-], 1 ♂, K. M. King leg., 5.vii.1950, 1 ♀, 5. vii.1957, 2 ♀♀, 9.vii.1957, 2 ♂♂, A. R. Brooks leg.; Sturgis, 51°56'N 102°32'W, 1.vii.1955, 1 ♂, J. R. Vockeroth leg. (all CNCI). YUKON: Klondike Hwy, 15 km S Carmacks, sweep sedges & rushes at pond margin, 11.vi.1997, 1 ♂ 3 ♀♀ ( LEMQ 0039972, -73, -94, 0040012); Klondike Hwy, S of Carmacks, Montague Road House, sweep roadside vegetation, 11.vi.1997, 3 ♀♀ ( LEMQ 0040008, -09, -13), all S. Boucher leg.; Dawson, 14.vi.1949, 2 ♂♂, 16.vi.1949, 1 ♂, 1.vii.1949, 3 ♀♀, W. W. Judd leg. ( CNCI); Emerald Lake, 60°15.8'N 134°45.1'W, sweep at lake margin, 20.vii.1998, 2 ♂♂ 1 ♀ ( LEMQ 0039934, -35, 0040017); Alaska Hwy at M’[C]lintock R., 60°33.6'N 134°29.4'W, sweep river edge, 14.vii.1998, 1 ♂ ( LEMQ 0039936); Tagish Campground, 60°18.8'N 134°16.1'W, sweep vegetation at campsite, 26.vi.1997, 3 ♂♂ ( LEMQ 0039931, -49, -50); Tagish Campground, 60°19'N 134°15'W, sweep vegetation at campsite, 26.vi.1997, 7 ♂♂ 6 ♀♀ ( LEMQ 0039933, -76–87); Tagish,Tagish Govt. Campground, 60°18'N 134°16'W, sweep grass at campsite, 26.vi.1997, 2 ♀♀ ( LEMQ 0039974, -75); Tagish, Tagish Govt. Campground, swept in open grassy area, 26.vi.1997, 1 ♂ 1 ♀ ( LEMQ 0040433, -34); Whitehorse, Bert Law Park at Yukon R., 60°42'N 135°02'W, sweep clearing in open forest, 27.vi.1997, 3 ♂♂ ( LEMQ 0039954, -69, -71); Whitehorse, Miles Canyon, 60°40'N 135°01.8'W, sweep grass/ sedges at Yukon River margin, 27.vii.1998, 3 ♂♂ 1 ♀ ( LEMQ 0039956, 0040003, -04, -16); Alaska Highway at Yarger Lake, sweep sedge & grasses along lake margin, 22.vi.1997, 1 ♂ ( LEMQ 0039968), all T. A. Wheeler leg.; Alaska Highway at Yukon River crossing, sweep sedges & grasses along river, 2.vii.1997, 1 ♂ 2 ♀♀, S. Boucher leg. ( LEMQ 0039937, -96, -97); same locality but 60°34'N 134°40'W, sweep grass/ sedges along river margin, 2.vii.1997, 2 ♂♂ 2 ♀♀, T. A. Wheeler leg. ( LEMQ 0039920, -21, -40, -41). UNITED STATES OF AMERICA: ALASKA: Fairbanks, [-]. viii.1960, 1 ♀, M. R. Wheeler & L. Throckmorton leg. ( AMNH); Fairbanks, Birch Hill, Alaska Ins[ect] Project, 15.viii.1948, 1 ♂, S. Lienk & C. O.Esselbaugh leg. ( USNM); Kenai Peninsula Borough, Long Term Ecological Monitoring Program, 3324, ~ 1 km N Crooked Ck. & ~1.5 km W Kolomin Lk., forest, sweep net sample, 20.vi.2004, 2 ♂♂ 2 ♀♀, Grimes leg. ( KNWR, 1 ♂ 1 ♀ ID:1819, 2118); Knik Lake, NW of Wasilla, sweeping vegetation, 18.vii.1978, 1 ♂ 1 ♀; same locality but sweeping vegetation edge of lake, 18.vii.1978, 2 ♂♂ 1 ♀, all P. H. Arnaud Jr. leg. (all CASC); Kodiak, [-]. vii.1960, 2 ♂♂, M. R. Wheeler leg. ( AMNH); Mat-Su, Eklutna (Knik Arm), 61°28.2'N 149°21.4'W, 7.viii.2002, 1 ♀, D. & W. N. Mathis leg. ( USNM); Naknek, on tundra, 21.vii.1952, 1 ♀, W.R. Mason leg. ( CNCI); Nenana, 16.vi.1953, 1 ♀, R. I. Sailer leg.; Ninilchik, 11.vii.1954, 1 ♂, R. Coleman leg. (both USNM); Tiekel River, Richardson Hwy ca. V-47, 1.viii.1978, 1 ♂, P. H. Arnaud Jr. leg. ( CASC); Tunnel Sta., 9.vi.1921, 1 ♀, J. M. Aldrich leg. ( USNM); Unalakleet, 18.vi.1961, 1 ♀, 19.vi.1961, 1 ♀, 6.vii.1961, 1 ♀, R. Madge leg., 22.vi.1961, 1 ♂, 5.vii.1961, 1 ♂, 8.vii.1961, 1 ♀, 12.vii.1961, 1 ♂, B. S. Heming leg. ( CNCI); Valdez, 5.vii.1948, 1 ♂, R. I. Sailer leg. ( USNM); Valdez, Valdez Glacier Campground, 2.viii.1978, 1 ♂, P. H. Arnaud Jr. leg. ( CASC). COLORADO: Clear Cr. Co., Chicago Cr., 8800', 25.viii.1961, 1 ♂, C. H. Mann leg. ( CNCI); Estes Park, 13.vii.1934, 1 ♀, A. L. Melander leg. ( USNM); Evans, Doolittle Ranch, 9800', 23.vii.1961, 1 ♂, J. G. Chillcott leg. ( CNCI); 3.5 mi S Florissant, Sanborn Ranch, swept from aspen grove nr. girls camp, 17.vii.2000, 2 ♂♂, 19.vii.2000, 1 ♂, B. A. Foote leg. ( CMNH); Summit Co., Frisco, 3.viii.2001, 1 ♂, I. S. Winkler leg. ( BYUC); Glen Haven, 14.viii.1952, 1 ♂, R. R. Dreisbach leg. ( USNM); Idaho Springs, 5 mi SW, 8600', 27.vii.1961, 2 ♂♂, J. G. Chillcott leg.; Loveland Pass, w. slope, 9850', 8.viii.1961, 1 ♂ (genit. prep.), B. H. Poole leg., 8.viii.1961, 1 ♀, C. H. Mann leg. (all CNCI); Boulder Co., Nederland, 24.vi.1961, 1 ♀, G. C. Steyskal leg. ( USNM); New Castle, 15.vii.1966, 2 ♀♀, W. Boyle & R. Lalonde leg. ( LEMQ); Rio Grande Co., South Fork, 8000', Malaise trap, 20.vi.1972, 2 ♀♀, W. W. Wirth leg. ( USNM); State Bridge nr. Bond, 7000', dry river bed and bank, 24–25.vi.1961, 2 ♀♀, C. H. Mann leg. ( CNCI); Eagle Co., State Bridge, Colorado R., 18.vi.1985, 1 ♀, D. W. Webb leg. ( INHS 40,208). IDAHO: Valley Co., Camp Cr., S. Fork Salmon River, 14.vi.1966, 1 ♂, L. S. Hawkins Jr. leg. ( WFBM); Franklin Co., Cub River Canyon, 2–11.vii.1985, 1 ♀, 7.viii.–17.ix.1985, 1 ♀, W.J. Hanson leg. ( LACM); Elmore Co., 6 mi E Featherville, 22.vi.1977, 1 ♀, [no collector]; Latah Co., Laird Park, 13 mi E Potlatch, 2800–3200', 4.vii.1983, 1 ♀, J. Jenkins leg.; Lemhi Co., 2 mi SW Lemhi, 9.vi.1976, 1 ♂, [no collector], 9.vi.1976, 1 ♂, M. W. Hanks leg. (all WFBM); Priest Lake, 1.viii.1916, 1 ♀, A. L. Melander leg. ( USNM); Custer Co., 6 mi E Stanley, 6200', 14.vii.1973, 1 ♂, Oman & Musgrave leg. ( OSAC). MAINE: Baxter St. Pk., bog nr. togue pond, 1–6.vii.1968, 1 ♀, D. M.Wood leg.( CNCI); Echo Lake, Mt.Desert, 12.vii.1918, 1 ♀, C.W. Johnson leg.; Seal Harbor, 29.vii.1930, 2 ♂♂, A. L. Melander leg.(all USNM). MICHIGAN: Houghton Co., 20.vi.1960, 1 ♂, R. & K. Dreisbach leg. ( USNM); Ironwood, 46°27.9'N 90°10.1'W, sweeps, low veg. Populus / Pinus , 22.vii.1999, 1 ♀; Ironwood, Mt. Zion Ski Area, 46°28.5'N 90°10.2'W, sweeps, low veg. in cut under Acer , 22.vii.1999, 1 ♂, both K. N. Barber leg. (both CNCI); Isle Royale, 3–7.viii.1936, 1 ♂, C. Sabrosky leg. ( USNM, S. W. Frost det. as A. tenuis ); Isle Royale, McCargo Cover, 10.vii.1966, 1 ♂, D. E. Bixler leg.; Keweenaw Co., Isle Royale, 10., 12.vii.1938, 1 ♀, G. Steyskal leg.; Mackinac Co., 4.vi.1959, 1 ♀, R. & K. Dreisbach leg. (all USNM). MINNESOTA: Eaglesnest, 13.vii.1957, 1 ♀ ( INHS 40,174), 30.vi.1959, 1 ♀ ( INHS 40,175), W. V. Balduf leg. MONTANA: 15 mi S Big Fork, 4.vii.1967, 1 ♂ 1 ♀, B. A. Foote leg. ( USNM); Gallatin Co., 79 mi S Bozeman, 7300', 11.vii.1973, 1 ♀, Oman & Musgrave leg. ( OSAC); Glacier Co., Duck Lake, 19.vii.1967, 1 ♂, D. Allen leg., 19.vii.1967, 1 ♂, B. A. Foote leg., 19.vii.1967, 2 ♂♂ 1 ♀, S. Whitney leg.; Saltese, 22.viii.1916, 1 ♂, A. L. Melander leg.; 33.0 mi N West Glacier, 5.vii.1966, 5 ♂♂, K. Valley leg. (all USNM). NEW HAMPSHIRE: “N.H.”, “Loew Coll.”,“ PARATYPE Anthomyza terminalis Loew No. 2122 ” (yellow label), “ Type 14558”, handwritten “ Anthomyza terminalis ” (red label), “ SYNTYPE CNC No.”, 1 ♂ (yellow label) ( CNCI, misidentined as a paratype of Anthophilina terminalis Loew ); “N.H.”, “178”, “Loew Coll.”, “ Type 14558”, 1 ♀ (red label) ( MCZC, misidentined as type specimen of Anthophilina terminalis Loew ); Colebrook, 30.v.1965, 1 ♂, D. M. Wood leg. ( CNCI); Franconia, 1 ♂, Mrs. Slosson leg. ( USNM, with det. as Anthomyza tenuis ); Gorham, 15.vii.1957, 1 ♀, J. R. Vockeroth leg. ( CNCI); Jefferson Notch, creek margin, 30.vii.1961, 1 ♂, W. W. Wirth leg.; Lost River, 7.vii.1931, 1 ♂, A. L. Melander leg. (both USNM); Mt. Washington, Auto Road, 3200', 14.viii.1958, 1 ♀, J. R. Vockeroth leg. ( CNCI); Mt. Washington, Dolly Copp, 4.vii.1936, 1 ♂ 1 ♀, 5.vii.1936, 1 ♂, A. L. Melander leg. ( USNM); Mt. Washington, Lakes of the Clouds, 5000', 31.viii.1954, 1 ♂, Becker, Munroe & Mason leg.; Coos Co., Pittsburg, 18.vi.1982, 1 ♀, J. R. Vockeroth leg. (both CNCI); Coos Co., 1 km E Stark, 44°36'N 71°24'W, sweep along Ammonoosuc R., 8.viii.2000, 1 ♂ 2 ♀♀ ( LEMQ 0039911, -18, -51); Coos Co., White Mt. National Forest, Dolly Copp, 44°19'N 71°13'W, sweep forest near river, 8.viii.2000, 1 ♀ ( LEMQ 0039953), all T. A. Wheeler leg.; Grafton Co., White Mt. Nat. Forest, Kancamagus Pass, 44°01'N 71°29'W, sweep, 10.viii.2000, 1 ♀, J. Forrest leg. ( LEMQ 0039952); Grafton Co., White Mtns. Nat. For., Long Pond, 675 m, 44°03'34.5"N 71°53'21.2"W, 30.vi.2004, 1 ♂, S. D. Gaimari leg. ( CSCA). NEW MEXICO: Santa Fe Co., 14 mi NE Santa Fe, 9600', aspen with stream, 18.vi.–3.vii.1979, 1 ♂ 1 ♀, S. & J. Peck leg. ( DEBU). NEW YORK: Adirondack Park, Blue Mountain, 43°52.5'N 74°25.8'W, sweep trail in mixed forest, 31.vii.2002, 1 ♂ 1 ♀ ( LEMQ 0040361, -62), sweep grasses at summit, 31.vii.2002, ♀ ( LEMQ 0040380), J. Forrest & T. A. Wheeler leg.; Adirondacks, Connery Pond, 15.vii.1938, 1 ♂; Adirondacks, Fish Creek, 2.viii.1929, 1 ♂, both A. L. Melander leg.; St. Lawrence Co., Cranberry Lk., swamp, 25.vi.1963, 2 ♂♂, W. W. Wirth leg.; Lake Placid, 28.vii.1929, 1 ♂, A. L. Melander leg. (all USNM); Lake Placid, 2000', 19.vii.1962, 3 ♀♀; Whiteface Mt., 19.vii.1962, 4600–4872', 1 ♂ 2 ♀♀, all J. R. Vockeroth leg.; Essex Co., Whiteface Mt., 44°21.96'N 73°54.18'W, ~ 1483 m, sweeps, mostly Calamagrostis at peak, 27.vii.2006, 15 ♂♂ 6 ♀♀; same locality but 44°22.16'N 73°53.88'W, ~ 1350 m, sweeps, graminoids, Rubus , 27.vii.2006, 1 ♂, all K. N.Barber leg. (all CNCI). OREGON: Benton Co., Mary’s Peak,~ 14.2 km WSW Flynn, 44°30.62'N 123°33.07'W, sweeps, sedge /herb meadow nr. parking, 10.vi.2009, 1 ♀, K. N. Barber leg. ( CNCI); Seaside, 26.vi.1963, 1 ♀, G. F. Knowlton leg. ( LACM); Tillamook Co., 1 mi W Sand Lake, 13.vi.1972, 1 ♂, W. N. Mathis leg. ( OSAC); Wallowa Lake, Hurricane Cr., 3.vii.1949, 2 ♂♂ 1 ♀ (2 ♂♂ on same pin), 11.vii.1949, 1 ♂ 1 ♀ (1 ♂ genit. prep.), J. L. & G. H. Sperry leg. ( USNM). SOUTH DAKOTA: Lawrence Co., #4, 6.2 mi SW Lead, 6000', 10.vii.1959, 1 ♀, G. W. Byers leg.; Pennington Co., Pactola Reservoir, 17.vi.1968, 1 ♀, “SLW” leg. (both OSAC). UTAH: Utah Co., Aspen Grove, [no date], 1 ♂, L. King leg. ( BYUC); Cache Co., Blacksmith Fork Canyon, 8–19.vii.1985, 1 ♀, 19.vi.–8. vii.1985, 1 ♂ 1 ♀, Malaise trap, 7–10.viii.1970, 1 ♂, W. J. Hanson leg.; Elk Park Creek, Uinta Mts., 17.vii.1952, 1 ♂, G. F. Knowlton & G. E. Bohart leg.; Cache Co., Green Canyon, 12–15.vi.1985, 1 ♀, 15–19.vi.1985, 1 ♀, 19–23. vi.1985, 2 ♂♂ 5 ♀♀, Malaise trap, 27–28.vi.1985, 1 ♀, N. Youssef leg. (all LACM); Cache Co., W Hodges Canyon, 23–27.vi.1978, 1 ♀, Hanson & Knowlton leg., 18–21.vii.1978, 1 ♀, Knowlton & Hanson leg.; Summit Co., 17 mi E Kamas, 5.viii.1971, 1 ♂, W. J. Hanson & G. F. Knowlton leg.; San Juan Co., La Sal Mtns., Geyser Pass, 10200', Malaise trap, 19.vii.1968, 1 ♂, [no collector]; Cache Co., Logan Canyon, 3.vii.1980, 1 ♀, Knowlton & Cazier leg., 8–15.ix.1993, 1 ♀, W. J. Hanson leg.; Cache Co., Logan Canyon, Bunchgrass Creek, 3–11.vii.1986, 1 ♂, Hanson & Knowlton leg.; same locality but Turner C.G. [campground], 16–30.viii.1985, 1 ♂, W. J. Hanson leg., 20.vi.1985, 1 ♀, G. F. Knowlton leg.; same locality but Twin Creek, 1–8.vii.1988, 1 ♀, 8–15.vii.1988, 3 ♀♀, 15–22.vii.1988, 1 ♂, W. J. Hanson leg. (all LACM); Utah Co., Mapleton, Malaise trap, 25.vi.–6.vii.1985, 1 ♀, [no collector]; San Juan Co., 10 mi SW Monticello, 20.vi.1971, 1 ♀, J. L Petty leg.; Cache Co., Tony Grove Cr., 12–18.viii.1995, 1 ♀, W. J. Hanson leg.; Cache Co., Tony Grove Jct., Malaise trap, 22–30.vi.1983, 2 ♀♀, 6–12.vii.1983, 1 ♀, 19–27. vii.1983, 1 ♂, 2–8.viii.1983, 1 ♂, 16–23.viii.1983, 1 ♂, 23–30.viii.1983, 1 ♂, 11–25.vii.1984, 4 ♀♀, 25–31.vii.1984, 1 ♂ 1 ♀, 1–13.viii.1984, 1 ♂, 13–21.viii.1984, 1 ♀, 1–7.ix.1984, 1 ♂, [no collector] (all LACM). WASHINGTON: Clallam Co., Bogachiel, 1.viii.1951, 2 ♂♂, M. R. Wheeler leg. ( AMNH); Pierce Co., Clover Creek, 9.vi.1979, 1 ♂, 15.vi.1979, 1 ♀, 7.vii.1979, 1 ♀, 20.vii.1979, 2 ♀♀, T. L. Whitworth leg. ( LACM); Longmire, 27.vi.1935, 5 ♂♂ 3 ♀♀; Mt. Constitution, 17.v.1910, 1 ♀, all A. L. Melander leg. (all USNM); Olympic N. P., 2 mi on Hoh Rain Forest Rd., 1797/1, 29.vi.1988, 1 ♀, J. A. Downes leg. ( CNCI); Olympic Nat. Park, Kalaloch, 8.vii.1968, 1 ♂ 1 ♀, W. W. Wirth leg. ( USNM); Pierce Co., Pleasant Valley, 1.viii.1977, 1 ♀, 4.viii.1978, 1 ♀, T. L. Whitworth leg. ( LACM); Seattle, 16.vi.1920, 1 ♂, A. L. Melander leg. ( USNM). WYOMING: Teton Co., 3 mi SW Hoback Jct., Teton Nat. For., 22.vi.1985, 1 ♂ 1 ♀, D. W. Webb leg. ( INHS 40,209, -210); Park Co., Pahaska Tepee, 6800', riverside meadow & pine forest, 24.viii.1979, 1 ♀, S. & J. Peck leg. ( DEBU); Yellowstone Pk.,Apollinaris, 8.vii.1923, 1 ♂; Yellowstone Pk., Spring Crk., 15.vii.1923, 1 ♂ 1 ♀; Yellowstone Pk., Thumb Sta., 16.vii.1923, 1 ♂; Yellowstone Pk., Twin Lakes, 10.vii.1923, 2 ♀♀; Yellowstone Pk., Upper Basin, 13.vii.1923, 1 ♀, all A. L. Melander leg. (all USNM).

Other material examined (not included in type series). CANADA: ALBERTA: Edmonton, University [of Alberta], 8.vi.1966, 1 ♂, K.A. Spencer leg. ( BMNH, abdomen lost) ; Edmonton, George Lake , 21.vi.1966, 1 ♀, K.A. Spencer leg. ( BMNH, damaged, one wing and head preserved in glycerine with genit. prep.) ; Jasper , 16.vi.1966, 1 ♂, K. A. Spencer leg. ( BMNH, headless) ; Peter Lougheed P. Pk. , 0.5 km W Little Highwood Pass, 50°38.55'N 115°02.94'W, sweeps, roadside grasses incl. Bromus inermis , 14.vii.2011, 1 ♀, K. N. Barber leg. ( DEBU 01502566 , teneral) . BRITISH COLUMBIA: Atlin , 2200', 7.vii.1955, 1 ♀, B. A. Gibbard leg. ( CNCI, headless) ; Crowsnest , Crowsnest Pass rest area, 11U 666201 5502396, 1377 m, ( CC 09-36 ), 17.viii.2009, 1 ♀, C. & D. Copley & J. Miskelly leg. ( RBCM ENT 009-006203 , poor shape, genit. prep.) ; Ft. Nelson , 13.vi.1948, 1 ♀, W. R. Mason leg. ( CNCI, greasy) ; Revelstoke , 2.vii.1973, 1 ♂, H. J. Teskey leg. ( CNCI, headless) . ONTARIO: Lake Superior P. Pk. , 47°16.42'N 84°33.66'W, pooter, grasses in roadside ditch, 15.vi.2004, 1 ♀, K. N. Barber leg. ( DEBU 01501453 , teneral) ; Lake Superior Prov. Park, Crescent Lake Trail , 47°16.81'N 84°33.23'W, sweeping, mostly graminoids under Betula / Acer , 9.vii.2010, 2 ♂♂ 1 ♀, J. Roháček leg. ( SMOC, 2 ♂♂ used for molecular work, 1 ♂ 1 ♀ headless) ; Thunder Bay , 22.vii.1970, 1 ♂, K. J. G. Deacon leg. ( DEBU, headless, genit. prep.) ; Waubamick [sic Waubamik], [-]. vi.1915, 1 ♂, H. S. Parish leg. ( USNM, headless) . SASKATCHEWAN: Kenosee , 7.vi.1958, 1 ♀, A. R. Brooks leg. ( CNCI, headless) . UNITED STATES OF AMERICA: ALASKA: Palmer , 24.vii.1958, 1 ♀, R. H.Washburn leg. ( UAMF 100022769 ) . COLORADO: Rocky Mt. N. P., 26.vii.1938, 1 ♀, H. H. Ross leg. ( INHS 40,180 , greasy, genit. prep.) . UTAH: Cache Co., Green Canyon , 19–23.vi.1985, 1 ♀, N. N. Youssef leg. ( LACM, poor condition) . WYOMING: Yellowstone Park, Imperial Geyser , 2.viii.1934, 1 ♀, A. L. Melander leg. ( USNM, abdomen missing) .

Description. Male. Total body length 2.18–2.94 mm; bicolourous, yellow ventrally and laterally and (relatively pale) brown dorsally, largely dull, dorsal parts of thorax and abdomen distinctly pale grey microtomentose. Head slightly higher than long, anteriorly (at antennal base) rather rounded in pronle with face very slightly receding, almost entirely yellow, only ocellar triangle and lateral spots on occiput brownish. Occiput slightly concave, medially with usual pair of silvery white microtomentose spots (largely fused ventrally above foramen), laterally to them at least with crescent-shaped ochreous to brown areas which, in darkest specimens, can be expanded dorsolaterally to reach eye margin. Frons relatively narrow, largely dull yellow, lighter anteriorly and on orbits, darker (to orange) medially in front of frontal triangle; ocellar triangle brown to dark brown and greyish microtomentose; frontal triangle dark yellow to orange and dull, narrowly margined by some golden microtomentum; stripes between the latter and orbits silky yellow; orbits whitish microtomentose up to posterior ors, subshining more posteriorly. Frontal triangle narrow and rather short, reaching at most to anterior third of frons. Frontal lunule minute, yellow. Face narrow, medially concave to folded, and, like parafacialia and gena, yellowish white with silvery white microtomentum; marginal stripe of parafacialia and gena golden yellow; postgena and mouthparts whitish yellow to yellow. Cephalic chaetotaxy (setae usually paler than in relatives): pvt relatively long and often strongly crossed; vti and oc longest of cephalic setae; vte and posterior ors usually slightly shorter than vti; 3 ors but only 2 long, middle ors slightly shorter than to as long as posterior ors; 1 shorter anterior ors setula well developed; 1–2 (usually) pairs of medial microsetulae in anterior third of frons but none in front of anterior ors setula; 1 small setula behind vte; postocular setulae (7–8) short, in single row; postgena with several setulae and 2 relatively small posteroventral setae; 1 vi (usually weaker but sometimes as long as posterior ors), subvibrissa small, normally slightly longer than anterior peristomal; only 3–4 nne peristomal setulae. Palpus light yellow, with 1 ventral preapical seta and several (6–7) small setulae subapically and ventrally. Eye convex, subovoid (widest in anteroventral half), with longest diameter oblique and about 1.4 times as long as shortest. Gena relatively high; its shortest height about 0.15 times as long as shortest eye diameter. Antenna strongly geniculate, pale yellow, 1st nagellomere with short white pilosity. Arista brown but basal segments yellow, about 1.8–1.9 times as long as antenna, with shorter ciliation than on 1st nagellomere.

Thorax slightly narrower than head, laterally yellow but dorsally mostly brown with light grey (often with bluish tinge) microtomentum. Scutum entirely or largely brown to greyish brown, in latter case with yellow or ochreous longitudinal vittae (of various length) on dc lines, in palest specimens also anterior third of scutum yellow; notopleural area and dorsal part of humeral callus yellow but lateral side of the latter always brownish; scutellum yellow on disc only in lightest specimens, otherwise partly or completely brown; both scutum and scutellum distinctly pale grey microtomentose and dull. Pleural part of thorax more shining than scutum, all brightly yellow, only in darkest specimens with very narrow brownish darkening at dorsal (rarely also posterior) margin of mesopleuron. Postscutellum and postnotum brown to dark brown. Thoracic chaetotaxy (setae usually paler than in relatives): 1 hu plus 2 (1 longer) setulae on humeral callus; 2 npl (anterior longer than hu, posterior shorter); 1 weak prs (about as long as hu); 1 sa and 1 pa (both weak as prs, pa often shorter); 2 usual postsutural dc (anterior markedly shorter) and 4–6 dc microsetae in front of them (the hindmost distinctly enlarged, sometimes resembling a 3rd dc macroseta); 3–4 rows of ac microsetae on suture, only 2 rows between dc macrosetae and posteriorly not reaching to level of posterior dc; 2 sc, apical as long as posterior dc (longest thoracic setae), laterobasal very weak, shorter than sa; 1 minute ppl; 2 long stpl, anterior shorter, and 2–4 setulae in dorsal half of sternopleuron, its ventral part with 3–4 longer setae. Scutellum rounded triangular, very slightly convex dorsally. Legs yellow to pale yellow, only last tarsal segments of all tarsi dark brown in distal half to three-fourths; setosity of all legs pale ochreous yellow except for brown ctenidial spine, ventroapical seta on t 2 and ventral setulae on mid basitarsus. f 1 with ctenidial spine slightly to distinctly longer than maximum width of t 1. f 3 with posteroventral row of sparse erect setae; only 4–5 of them in distal two-nfths shortened and thickened; t 2 with moderate ventroapical seta; fore and hind basitarsus proximoventrally with several (usually 2–3) longer pale setulae; also 1–2 proximoventral setulae in mid basitarsus somewhat prolonged but brown; other parts of legs simply setulose. Wing ( Fig. 274 ) elongate, moderately narrow, with pale yellowish ochreous veins and membrane. C with very sparse spinulae between apices of R 1 and R 2+3. R 2+3 long, parallel to C with apex very slightly upcurving; R 4+5 very slightly bent to almost straight, usually very slightly divergent from M (this almost straight) towards apex. Discal cell (dm) long, moderately narrow; r-m situated near the middle of cell dm or slightly basal to the midpoint. Apical portion of CuA 1 slightly to distinctly longer than dm-cu and almost reaching wing margin; A 1 short, ending far from it. Alula distinct but narrow. Wing measurements: length 2.18–2.88 mm, width 0.73–0.99 mm, Cs 3: Cs 4 = 1.03–1.32, rm\dm-cu: dm-cu = 1.82–2.65. Haltere pale yellow to yellowish white.

Abdomen. Preabdominal terga pale brown to brown, subshining but distinctly pale grey microtomentose and relatively shortly and usually pale setose. T1 and T2 only laterally partly fused; T1 shorter than T2 and distinctly narrower than T3. T3–T5 subequal in size, reaching onto lateroventral side of abdomen. Preabdominal sterna pale yellow (S2 sometimes darker yellow), not very narrow and becoming wider posteriorly, S2 slightly, S3–S4 more and S5 distinctly transversely trapezoidal, the latter being the largest and widest sternum; sterna nnely setose, only S1 bare and with darker posterior marginal stripe. T6 short (less than one-third of T5), transverse, bare, medially broadly unpigmented, with only lateral parts pale brown. S6 and S7 strongly asymmetrical and fused dorsally, brown as T5, both with more sclerotized and dark brown anterior margin; S6 with 2–3, S7 with 2–4 distinct setae; S8 slightly longer than epandrium, tapered posteriorly, often somewhat darker than S6 and S7 and setose in posterior half.

Genitalia. Epandrium ( Figs 260, 261 , 294 ) brown to dark brown, concolourous with or somewhat darker than pregenital sclerites in contrast to that of A. pallida , relatively long and broad (longer and wider than in A. pallida ) and moderately setose, with 2 pairs of longer and thicker dorsomedial setae; also anal nssure larger and wider than in latter species. Cercus rather short and broad, with numerous nne setae, apical longest. Medandrium ( Fig. 260 ) distinctly wider and with more projecting dorsolateral corners than in A. pallida . Gonostylus ( Figs 260, 261, 263 ) slightly incurved, largely micropubescent on outer side and sparsely setose on inner side, as in A. pallida , but shape relatively short and oblong (not tapered and triangular), and distally with 3 distinct teeth (1 apical and 2 subapical). Hypandrium ( Fig. 262 ) also similar to that of A. pallida , being medium-sized, relatively slender and with anterior internal lobes reduced. Transandrium ( Fig. 265 ) simple, slightly bent medially and without caudal process as in A. pallida , but somewhat broader. Pregonite ( Fig. 262 ) strongly resembling that of A. pallida , having a small, short, triangular process behind the larger anterior tooth-like projection, but the latter process is trapezoidal rather than subtriangular, and the posterior part of the pregonite has more (5–6) setae. Postgonite ( Fig. 262 ) relatively robust, less widened distally compared to A. pallida , with 1 anterior setula in proximal fourth, several sensillae and blunt apex; its proximal part connected with slender basal sclerite. Dorsal internal sclerite at base of postgonite narrow but distinct. Basal membrane ( Figs 262, 265 ) densely overgrown with pigmented and distinctly larger spines than those in A. pallida . Aedeagal part of folding apparatus with a group (larger than in A. pallida ) of grain-like tubercles and usual elongate hyaline warts ( Fig. 264 ). Connecting sclerite ( Fig. 264 ) slender, dark-pigmented and, in contrast to that of A. pallida , terminating in a group of distinct pigmented spines. Phallapodeme ( Fig. 264 ) relatively slender, with basal part deeply forked; fulcrum shifted close to apex as in A. pallida , but apex more projecting and wing-like laterally. Aedeagus with short phallophore ( Fig. 264 ) and very large and long distiphallus that is bind from near its base. Saccus unusually long and proximally distinctly sclerotized as in A.pallida , but its large membranous part is armed with 4 robust dark spines. Filum slender, long and arched, very similarly constructed to that of A. pallida , but basally broader and partly micropubescent, medially curved and with membranous fringe, and distally with distinct sharp dorsal tooth and thin, curved, acutely pointed apex ( Fig. 266 ), the latter provided with 3 subterminal spinulae. Ejacapodeme very small and pale-pigmented, with minute and pointed terminal projection ( Fig. 264 ).

Female. Similar to male unless mentioned otherwise. Total body length 2.58–3.30 mm. Generally paler with brown markings reduced. Antenna with outer side of 1st nagellomere distinctly darkened, ochreous orange to pale brown, with only margins variably yellow. Darkened pattern on scutum and scutellum more reduced, appearing as longitudially greyish brown-and-yellow stripes, sometimes even with yellow areas prevailing. f 3 simply setulose posteroventrally. Wing measurements: length 2.42–3.41 mm, width 0.81–1.13 mm, Cs 3: Cs 4 = 0.97–1.26, rm\dm-cu: dm-cu = 2.07–2.84. Abdomen with preabdominal terga variable in colour, ranging from almost entirely yellow (only T1 with medial pale brown darkening) to proximally (T1, T2) completely pale brown to brown and distally (T3–T5) with large medial brownish areas. T2–T5 shorter and more transverse than in male. Preabdominal sterna hardly shorter than in male, similarly coloured but rather transversely suboblong than trapezoidal. S2–S3 slightly, S4–S5 distinctly transverse, S5 largest.

Postabdomen ( Figs 268–270 ) relatively long and narrow, tapered posteriorly, telescopic, with sclerites yellow and/or (partly) brown. T6 simple, narrower than T5 (longer than in A. pallida ), trapezoidal, entirely yellow or variably brown medially ( Fig. 268 ) and with relatively short setae. S6 narrower than S5 (and less transverse compared to A. pallida ), transversely suboblong, nnely setose. T7 slightly tapered posteriorly, always (in contrast to that of A. pallida ) with large brown area of various length posteromedially, yellow laterally and anteriorly, setae relatively short and nne, and its sides expanded onto venter but not with anterior corners touching ( Fig. 270 ) and not fused to S7; S7 ( Fig. 270 ) small and narrow, widened posteriorly and nnely setose. 7th spiracle embedded in T7 far from anteroventrolateral margin ( Fig. 269 ). 8th segment densely micropubescent laterally. T8 brown, distinctly longer than broad, anteriorly slightly tapered and deeply emarginate ( Fig. 268 ), with nne exclinate setae and reduced micropubescence; S8 slightly shorter than T8, medially divided into 2 sclerites (together of cordate shape, Fig. 270 ) each with 2 longer nne setae besides short setosity and with posterior end invaginated into 8th segment. Genital chamber (uterus) with internal sclerotization well developed ( Figs 272, 273 ) although rather pale-pigmented, formed by 2 separate pairs of nat curved sclerites (anterior pair higher, posterior pair longer, hence different from those in A. pallida , where subequal, cf. ROHÁĆEK 2006a: Figs 237, 238 ) and 1 anterior, slender, strongly curved annular sclerite ( Fig. 273 ). Distal part of genital chamber often with dark secondary spine-like excrescences ( Fig. 273 ). Ventral receptacle ( Fig. 273 ) slender, tube-like, long, hyaline, terminally spirally twisted, proximally broader but without subbasal attenuation seen in A. pallida ; accessory gland hyaline, with minute granulae on surface, borne on distally dilated and distinctly ringed duct. Spermathecae (1+1) shortly ovoid ( Figs 267, 271 ), each with short cup-like terminal invagination (wider and shorter than in A. pallida ), very nnely spinulose base and smooth surface (not nnely ringed in distal half as that of A. pallida ); spermathecal duct very long, terminally without distinct cervix. T10 small, darker than in A. pallida and of subcordate shape, with 1 medial pair of long setae and very reduced micropubescence. S10 also different from that of A. pallida , elongately suboblong thus narrow and longer, nnely setulose and micropubescent. Cerci moderately sized, separated by their width ( Fig. 268 ); with numerous nne setae, apical and dorsopreapical longest.

Discussion. This new species, Anthomyza pengellyi sp. nov., belongs to the A. pallida group as denned above. It is distinctly different from all other Nearctic relatives of the A. pallida group (see the key) and is, in fact, closest to the Palaearctic A. pallida forming with it the A. pallida (s. str.) subgroup. Their sister-species relationship is demonstrated by general resemblance (including the brown lateral side of the humeral callus) and, more importantly, by several shared synapomorphies that denne the subgroup: gonostylus shortened; pregonite with broadened anterior tooth and short posterior triangular process; saccus of distiphallus prolonged; nlum of distiphallus with characteristic subterminal tooth; and internal pairs of sclerites of female genital chamber situated as anterior and posterior (not dorsal and ventral). The most distinct differences between A. pengellyi and A. pallida can be seen in the structures of the male genitalia (shape of gonostylus, armature of basal membrane, saccus and connecting sclerite) and female postabdomen (shape and/or pigmentation of T6, S6, T7, T8, T10, structure of spermatheca, form of internal sclerites of female genital chamber) but A. pengellyi is most readily differentiated by the distinct brown to greyish brown markings on the mesonotum (largely yellow or with very faint marking in A. pallida ), a brown to dark brown subscutellum and postnotum (reddish brown to ochreous in A. pallida ), a brown epandrium (yellow in A. pallida ), and often by pale setae that are ochreous to yellow in most specimens but ranging to black (always brown to black in A. pallida ).

Etymology. This species is dedicated to the memory of Professor David Pengelly who formally introduced the junior author, as an undergraduate and graduate student, to the wonders of the insect world.

Biology. The habitat for A. pengellyi is still rather ill-denned but graminoids are the most common component. Collections have often been made from dry sites dominated by grasses to moist sites dominated by sedges and various combinations of graminoids. Dry (open disturbed) sites with grasses have included Bromus inermis , Poa compressa L., and Leymus innovatus (Beal) Pilg. in the west (Alberta: Peter Lougheed P. Pk. – 0.5 km W Little Highwood Pass, Fig. 259 ; ~ 26 km NW Highwood House – Mt. Lipsett pulloff; Elbow-Sheep Wildland Pk. – ~22.5 km NW Highwood House) while a much less disturbed site under Betula / Acer had a dominant graminoid component in the east (Ontario: Lake Superior Prov. Park – Crescent Lake Trail, Fig. 258 ). A decidedly mesic site with mostly Glyceria striata (Lam.) Hitchc. (Ontario: ~ 61 km NNW Elliot Lake – Three Lakes) was shared with A. dichroa and A. mcalpinei , while an extremely xeric sandy beach supporting the beachgrass Ammophila breviligulata Fernald (Ontario: Pancake Bay P. Pk., Fig. 546 ), was shared with the more numerous A. vulgaris . Additional references are made to “grass slope” and “ grasses at edge of agricultural neld” (Alberta: Dunvegan) which sound similar to the “streamside, mostly grasses ” reference (Alberta: ~ 20 km SSE Robb). More specincally, Calamagrostis sp. (probably C. canadensis ) was identined in a dry site (New York: Whiteface Mt.) and C. canadensis in a wet hydro cut, either by itself or mixed with Equisetum ssuviatile (Ontario: Otter Rapids).

A mesic forested site yielded A. pengellyi from a range of “trailside graminoids, Impatiens ”, “short trail grasses ”, “trailside grasses ” and “trailside Carex ” (Ontario: Sault Ste. Marie – Sault College Outdoor Lab). The wet border of a fen habitat dominated by a mix of Carex utriculata Boott in Hook. and Poa cf. pratensis L. also yielded A. pengellyi , a site that transitioned to a sparse growth of mixed herbs and grasses (probably including P. cf. pratensis ) near the roadside (Alberta: Peter Lougheed P. Pk). But a similarly dry border of a fen with sparse growth of C. utriculata , without a strong grass component, was also productive (Alberta: Peter Lougheed P. Pk. – Upper Lake Drive). Another example where graminoids were either absent or an insignincant component was of a community dominated by Eurybia and Equisetum under an aspen canopy (Ontario: Otter Rapids). The edge of a wet forest trail (Ontario: Moosonee) dominated by Equisetum , Rubus , and Cornus but with a signincant graminoid component was also a good site for A. pengellyi . The night period for A. pengellyi runs from 26 May (Alberta: Elkwater Park; Quebec: Old Chelsea) to 8–15 September (Utah: Logan Canyon).

Distribution. Anthomyza pengellyi has a broad transcontinental distribution from Labrador to Nova Scotia, Maine, New York, Michigan and Minnesota, west to Northwest Territories, Alaska and Oregon, and south to Utah, Colorado and New Mexico. Records are available for Canada: Alberta, British Columbia, Labrador, Manitoba, New Brunswick, Northwest Territories, Nova Scotia, Ontario, Quebec, Saskatchewan, Yukon; United States of America: Alaska, Colorado, Idaho, Maine, Michigan, Minnesota, Montana, New Hampshire, New Mexico, New York, Oregon, South Dakota, Utah, Washington, Wyoming (see Table 2).

Figures

Figs 256–259. Living Anthomyza pengellyi sp. nov. and its habitats. 256 – A. pengellyi sp. nov., male, laterally, body length ca. 2.5 mm; 257 – female, subdorsally, body length ca. 2.9 mm (both Canada: Ontario: Lake Superior Prov. Park – Crescent Lake Trail); 258 – grassy undergrowth of birch-maple forest at Crescent Lake (Canada: Ontario), natural habitat of A. pengellyi; 259 – roadside grasses with predominant Bromus inermis near Little Highwood Pass (Canada:Alberta), secondary habitat of A. pengellyi. Photo by J.Roháček (Figs 256–258) and K. N. Barber (Fig.259).

Figs 260–266. Anthomyza pengellyi sp. nov., paratype male (Canada: Ontario). 260 – external genitalia, caudally; 261 – the same, laterally; 262 – hypandrial complex, laterally; 263 – gonostylus, laterocaudally (widest extension); 264 – aedeagal complex, laterally; 265 – transandrium,caudally; 266 – apex of nlum, sublaterally (widest extension). Scales = 0.05 mm (Figs 263, 266) and 0.1 mm (others).

Figs 267–273. Anthomyza pengellyi sp. nov., paratype female (Canada: Ontario). 267 – spermatheca; 268 – postabdomen, dorsally; 269 – the same, laterally; 270 – the same, ventrally; 271 – spermatheca; 272 – female internal sclerites, ventrally; 273 – female genital chamber laterally.Scales = 0.05 mm (Figs 267, 271), 0.2 mm (Figs 268–270) and 0.1 mm (others). For abbreviations see p. 11.

Figs 274–276. Wings of the Nearctic species of the Anthomyza pallida group. 274 – A. pengellyi sp. nov., male, wing length 2.65 mm (Canada: Ontario); 275 – A. mcalpinei sp. nov., male, wing length 2.7 mm (Canada: Ontario); 276 – A. pullinotum sp. nov., male, wing length 2.3 mm (Canada: Yukon Territory). Photo by K. N. Barber.

Figs 294–298. Primary types of the Anthomyza pallida group. 294 – A. pengellyi sp. nov, holotype male, laterally, body length 2.44 mm; 295 – A.mcalpinei sp. nov., holotype male, laterally, body length 2.94 mm; 296 – A. pullinotum sp. nov., holotype male, laterally, body length 2.74 mm; 297 – the same, head and thorax, laterally; 298 – the same, abdomen laterally. Photo by J. Roháček.

Figs 232–238. Anthomyza oblonga sp. nov., paratype female (Canada: Ontario). 232 – abdomen, dorsally; 233 –female genital chamber and S8 (with setosity omitted), laterally; 234 – postabdomen, dorsally; 235 – spermatheca; 236 – internal sclerites, ventrally; 237 – spermatheca; 238 – postabdomen, ventrally. Scales = 0.3 mm (Fig. 232), 0.05 mm (Figs 235, 237) and 0.1 mm (others).

Figs 544–546. Habitats of Nearctic species of the Anthomyza gracilis group and A. vulgaris sp. nov. 544 – riverbank of Mississagi River (Canada: Ontario) with rich graminoid communities, habitat in the type locality of A. furvifrons sp. nov.; 545 – A. vulgaris sp. nov., female on leaf of sedge, body length ca. 2.8 mm (Canada: Ontario: Sault Ste Marie); 546 – sandy beach with growth of Ammophila breviligulata (Canada:Ontario: Pancake Bay), extreme habitat of A. vulgaris. Photo by K. N. Barber (Fig. 544) and J. Roháček (others).