HOLOPSENELLINAE (MODIFIED FROM ENGEL, 2019 )

Jouault, Corentin, Ngô-Muller, Valérie, Pouillon, Jean-Marc & Nel, André, 2021, New Burmese amber fossils clarify the evolution of bethylid wasps (Hymenoptera: Chrysidoidea), Zoological Journal of the Linnean Society 191 (4), pp. 1044-1058 : 1051

KEY TO GENERA OF † HOLOPSENELLINAE (MODIFIED FROM ENGEL, 2019) 1. Clypeus with apical margin unmodified; first metasomal tergum without transverse carina .....................2

- Clypeus with apical margin modified and medially emarginate; first metasomal tergum with transverse carina sometimes present [Early Cretaceous (Barremian): Lebanon] ...........................................................3

2. Anterior wing margin beyond pterostigma swollen (margin discontinuous); second cubital cell open; pretarsal claws simple (Cenomanian: Myanmar).............................................. Holopsenelliscus Engel, 2019

- Anterior wing margin beyond pterostigma not swollen (margin continuous); second cubital cell closed; pretarsal claws dentate (Santonian: Taimyr amber, Russia) .................................. Cretabythus Evans, 1973

3. Antennae with ten flagellomeres; forewings with cell 2R1 short, ending far from wing apex; veins Rs and 2r-rs curved; propodeum with propodeal spines (Cenomanian: Myanmar) ................... Holopsenella Engel et al., 2016

- Antennae with 11 flagellomeres; forewings with cell 2R1 elongate, nearly reaching wing apex; veins Rs and 2r-rs straight; propodeum without propodeal spines (Cenomanian: Myanmar) ...... Megalopsenella gen. nov.