Speonemadus vandalitiae ( Heyden, 1870 )

Fresneda, Javier, Faille, Arnaud, Fery, Hans & Ribera, Ignacio, 2019, A molecular phylogeny of Speonemadus Jeannel, 1922 with description of two new species from Morocco (Coleoptera: Leiodidae: Cholevinae: Anemadini), Zootaxa 4543 (1), pp. 1-36 : 20-22

Speonemadus vandalitiae ( Heyden, 1870)

Figs 10 , 17 , 37

Catops vandalitiae Heyden, 1870: 97 .

Ptomaphagus vandalitiae ( Heyden); Heyden 1883 : 368.

Anemadus vandalitiae (Heyden) : Uhagón 1890: 32.

Hormosacus vandalitiae (Heyden) ; Jeannel 1936: 214.

Hormosacus vandalitiae pyrenaeus Jeannel (1936: 214) ; Blas 1984: 252 (synonymy)

Hormosacus vandalitiae pyrenaeus Jeannel ;

Speonemadus vandalitiae (Heyden) ; Giachino & Vailati 1993: 183.

Type locality. "Sierra della Nieve prope Ronda Andalusiae" ( Heyden 1870), Spain.

Lectotype ( Fig. 37 ): ♂, "Ronda, Heyden" [hw L. von Heyden], " Syntypus " [red, printed], microscopic slide with aedeagus and "1563" [hw], " Coll. L. v. Heyden, DEI Eberswalde" [printed], " Lectotypus ♂, Catops Vandalitiae Heyden, P. M. Giachino & D. Vailati des. 1992" [red, printed] ( SDEI).

Diagnosis. Apex of median lobe acuminated ( Fig. 10 ); parameres very long, forming a marked angle, with apex round and expansion on inner side ( Fig. 17 ) (see other characters under S. gracilis , see above). Molecular data place S. vandalitiae as sister species of S. pulchellus ; although both species belong to the vandalitiae -group sensu Giachino & Vailati (1993), their aedeagi are considerably different (compare Figs 10 and 11 ), with much shorter and robust parameres relative to the length of the median lobe in S. pulchellus .

Distribution ( Fig. 39 ; Appendix). Iberian Peninsula, extending to the French Pyrenees ( Jeannel 1936; Giachino & Vailati 1993; Perreau 2000, 2004; Salgado et al. 2008; Agirre & Blas 2009; Fresneda & Salgado 2017). In Spain there are records from Albacete, Ciudad Real, Granada, Huesca and Madrid ( Fresneda et al. 2007), Málaga ( Heyden 1870; Fuente 1925; Giachino & Vailati 1993), Murcia ( Fresneda et al. 2007), Tarragona and Toledo; and in France from Basses-Pyrénées ( Jeannel 1936). However, some of these records may refer to S. gracilis , which has been considered to be a synonym of S. vandalitiae since Reitter (1885) (see section 3.3. above).

Figures

FIGURE 37. Lectotype of Catops vandalitiae (scale bar: specimen, 2 mm; aedeagus, 0.3 mm).

FIGURE 39. Distribution maps of: (39) S. gracilis, S. vandalitiae. See Appendix for detailed locality data.

FIGURES 10–11. Aedeagi in ventral view of: (10) S. vandalitiae (Villanúa, Spain); (11) S. pulchellus (Ficuzza, Sicily).

FIGURES 13–28. Apex of the parameres of: (13) Speonemadus orchesioides (Ammi-Moussa, Algeria); (14) holotype of S. brusteli sp. n. (Talembote, Morocco); (15) S. gracilis (Hornos, Spain); (16) S. tenuipes (Ifri Semedane, Algeria); (17) S. vandalitiae (Villanúa, Spain); (18) S. pulchellus (Ficuzza, Sicily); (19) holotype of S. algarvensis (Gruta do Vale Telheiro, Portugal); (20) S. angusticollis (Rute, Spain); (21) S. bolivari (Ardales, Spain); (22) S. breuili (Motillas, Spain); (23) S. escalerai (Jumilla, Spain); (24) S. transversostriatus (La Bastida, Spain); (25) S. clathratus (Linarejos, Spain); (26) S. maroccanus (Taza, Morocco); (27) holotype of S. comasi sp. n. (Tagelft, Morocco); (28) Anemadus hajeki (Yunnan, China).