Speonemadus comasi,

Fresneda, Javier, Faille, Arnaud, Fery, Hans & Ribera, Ignacio, 2019, A molecular phylogeny of Speonemadus Jeannel, 1922 with description of two new species from Morocco (Coleoptera: Leiodidae: Cholevinae: Anemadini), Zootaxa 4543 (1), pp. 1-36: 6-9

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Speonemadus comasi

sp. n.

Speonemadus comasi  sp. n.

Figs 2View FIGURES 1–2, 3View FIGURES 3–4, 12, 27

Type locality: Morocco, Tagelft , Iri Adar Nyighil, 32°10'N 5°57'W, 2250 m ( Fig. 45View FIGURE 45)GoogleMaps  .

Holotype: ♂, "Iri Adar Nyighil / -Tagelft- Marroc / 9-VII-2005 / Fadrique- [hw] / Auroux-Comas Leg." [printed], " HOLOTYPUS / Speonemadus  / comasi  sp. n. / JF-AF-HF-IR det. 2017" [red label, hw] ( MZB). 

Description of the holotype: Habitus as in Fig. 2View FIGURES 1–2. Body length 3.6 mm. Body shape elongate, colour brownreddish, darker on head and pronotal disk. Head with eyes well developed, with visible suture between epistome and frons; surface with coarse, dense punctation; antennomeres VII–XI forming a loose club, antennomere X as long as wide. Pronotum transverse (maximum width 1.3 mm; maximum length 0.8 mm), sides strongly arched, still more strongly posteriorly; base of pronotum sinuate, as wide as base of elytra; surface with coarse, dense and rough punctation. Elytra very elongate, wider in basal third, regularly tapered towards apex; with parasutural stria; surface with fine and sparse transverse striolae. Legs with pentamerous tarsi; protibiae sinuate, more strongly on inner side ( Fig. 3View FIGURES 3–4); mesotibiae less sinuate, more regularly curved; first three tarsomeres of protarsi and first two tarsomeres of mesotarsi dilated. Male genitalia (Fig. 12). Basal lamina of aedeagus as long as apical part; parameres about as long as median lobe; median lobe in dorsal view with sides slightly sinuate, regularly tapering towards apex; apex with small, rounded extension; basal 2/3 of parameres straight, apical third with external side strongly dilated, narrowing towards apex; apex with triangular expansion on external side, with four fine setae and without conical spine ( Fig. 27View FIGURES 13–28); endophallus entirely covered with scales and spines, with two dark parallel longitudinal bands scarcely noticeable.

Females unknown.

Etymology. The species is dedicated to our friend and colleague Jordi Comas ( Spain, Barcelona), one of the collectors of the only known specimen of this most interesting species. The specific name is a substantive in the genitive case.

Remarks. Speonemadus comasi  sp. n. is a peculiar species, with characters similar to both Speonemadus  and Anemadus  . The general slender habitus and the modified protibiae are similar to the species of the escalerai  -group (e.g. S. angusticollis  , Fig. 4View FIGURES 3–4), and the shape of the parameres with their triangular apical expansion is similar to that of S. brusteli  sp. n. of the vandalitiae  -group. However, the endophallus is similar to that of the species of Anemadus  , without well defined groups of spines, such as those to the loebli  -group (e.g. A. loebli Giachino & Vailati, 1993  , from Turkey) or the strigosus  -group (e.g. A. strigosus (Kraatz, 1852)  from Czech Republic, Austria, Italy, Romania, Croatia, Bosnia-Herzegovina, Montenegro and Greece, A. bianchii Reitter, 1906  from Italy or A. arcadius Reitter, 1885  from Greece). In addition to the morphological characters, the distribution of the species supports its inclusion in Speonemadus  , as species of Anemadus  are found from Italy (including Sicily) and southern France to the East, reaching China ( Perreau 2004). In fact, this is one of the westernmost species of Speonemadus  , with other species of the genus distributed further east except for some dubious records of S. tenuipes  in the High Atlas (see below).

Distribution ( Fig. 38View FIGURE 38). So far only known from the type locality in Morocco.


Museum Zoologicum Bogoriense