Hadronotus insulare (Talamas, Bremer, Moore, Bon, Lahey, Roberts, Combee, McGathey, van Noort, Timokhov, Hougardy & Hogg, 2021)
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publication ID |
https://doi.org/ 10.1080/00222933.2024.2410028 |
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persistent identifier |
https://treatment.plazi.org/id/03FC879B-FF98-FFC1-9B80-FF3C81EB04F4 |
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treatment provided by |
Plazi |
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scientific name |
Hadronotus insulare |
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Key to females of the insulare species group of Hadronotus in the Oriental region
1. Basal foveae on T1 absent; T1 smooth with setigerous punctae without longitudinal costae; T2 smooth with setigerous punctae; entire metasoma covered with long setae ( Figures 9 View Figure 9 (a), 26(h)); frontal depression with oblique carinae on either side of central keel and predominantly smooth with large reticulations lateroventrally; IOS> 1.8× eye length; OOL> 1.3× MOD ( Figure 9 View Figure 9 (f)) .................................. H. gowatyae sp. n.
- Basal foveae on T1 present; T1 longitudinally costate; T2 never smooth, coarsely sculptured; metasoma partially covered with setae (eg Figure 26 View Figure 26 (c–f)); frontal depression transversely carinate, with narrow smooth areas between carinae and with polygonal cells or reticulations lateroventrally; IOS ≤ 1.5× eye length; OOL at most subequal to MOD (eg Figures 1 View Figure 1 (f), 4(f), 5(f), 7(f)) .................................................................... 2
2. Marginalis very short, indicated as a spot ( Figures 4 View Figure 4 (e), 7(e), 12(a), 19(d)); frontal depression triangular; submedian carina of frontal depression complete ( Figures 3 View Figure 3 (e), 4(f), 7(f), 12(e), 19(b)); mesopleuron predominantly transversely carinate ( Figures 4 View Figure 4 (c), 7(c), 12(c), 19(e)); body size generally small ...................................................................... 3
- Marginalis elongate (eg Figures 1 View Figure 1 (e), 5(e), 8(e), 10(e), 22(e)); frontal depression generally ovoid (trigonal in H. divergens sp. n.); submedian carina of frontal depression incomplete (eg Figures 1 View Figure 1 (f), 5(f), 8(f), 10(f), 22(f)); mesopleuron with depressions or foveae interspersed with transverse carinae (except in H. samantara , where it is entirely transversely carinate) (eg Figures 1 View Figure 1 (c), 5(c), 8(c), 10(c), 22(c)); body size generally large ...................................................................................................................................... 5
3. Postmarginalis present; submarginalis of fore wing parallel to anterior margin of wing before reaching marginalis; fore wing at least 2.4× as long as wide ( Figures 12 View Figure 12 (a), 19 (d)); in dorsal view metascutellum bulging as a blunt tubercle and projecting over propodeum; propodeum without a medial notch ( Figures 12 View Figure 12 (f), 19(a)); metasoma longer than wide ( Figures 26 View Figure 26 (k), 27(b)) ............................................................................................. ........................................................................ H. triatomae Masner and H. linshcostei Masner
- Postmarginalis absent; submarginalis of fore wing with a dip before reaching marginalis; fore wing at most 2× as long as wide ( Figures 4 View Figure 4 (e), 7(e)); metascutellum wide and not bulging, anteriorly foveate and posteriorly smooth, not projecting over propodeum; propodeum with a medial notch ( Figures 4 View Figure 4 (g), 7(g)); metasoma either as long as wide or wider than long ( Figure 26 View Figure 26 (c,f)) ............................................................................... 4
4. Central keel restricted to lower half of frontal depression; frontal depression wider than IOS, as inner orbits converge dorsally ( Figure 7 View Figure 7 (f)); anterior margin of T1 weakly convex; space between longitudinal costae on T1 smooth medially; propodeum deeply notched medially ( Figure 26 View Figure 26 (f)); transverse carinae on mesopleuron closely placed and space between them smooth ( Figure 7 View Figure 7 (c)); submarginalis dips weakly before reaching marginalis ( Figure 7 View Figure 7 (e)) ...................... ...................... H. galdikasae sp. n.
- Central keel extending the entire length of frontal depression with the keel projecting vertically; frontal depression as wide as IOS ( Figures 3 View Figure 3 (e), 4(f)); anterior margin of T1 projecting medially and laterally; space between longitudinal costae on T1 with rugose sculpture; propodeum with a shallow notch medially ( Figure 26 View Figure 26 (c)); transverse carinae of mesopleuron spaced apart and space between them with rugose sculpture ( Figure 4 View Figure 4 (c)); submarginalis dips extensively before reaching marginalis ( Figure 4 View Figure 4 (e)) ....................................................................................................................... H. circus (Kozlov and Lê)
5. Vertex sharp; vertex (in frontal view) above dorsal margin of compound eye ( Figure 16 View Figure 16 (d)); temples absent ( Figure 16 View Figure 16 (b)); metasoma as long as wide ( Figure 26 View Figure 26 (o)) ................. ................................................................................................................ H. strongist (Kozlov and Lê)
- Vertex not sharp; vertex in (frontal view) not raised above dorsal margin of compound eye, evenly rounded; temples distinct (eg Figures 1 View Figure 1 (g), 5(g), 6(g), 8(g)); metasoma longer than wide (eg Figure 26 View Figure 26 (a,d,g,l)) ..................................................................................... 6
6. Inner orbits on frons diverging dorsally in frontal view; frontal depression dorsally pointed ( Figure 6 View Figure 6 (f)); metapleuron entirely densely setose ( Figure 6 View Figure 6 (c)); basal foveae on T2 present; posterior tergites narrowing abruptly ( Figure 26 View Figure 26 (e)); entire lateral propodeal area setose ( Figure 6 View Figure 6 (g)) ............................ ............................ H. divergens sp. n.
- Inner orbits on frons either converging dorsally or parallel to each other in frontal view; frontal depression dorsally arched (eg Figures 1 View Figure 1 (f), 5(f), 8(f), 10(f), 22(f)); metapleuron partially setose, setae either dense or sparse (eg Figures 1 View Figure 1 (c), 5(c), 8(c), 10(c), 22(c)); basal foveae on T2 absent; posterior tergites narrowing gradually (eg Figures 26 View Figure 26 (a,d,g,i), 27(d)); lateral propodeal area generally without setae, if setae present, restricted to posterior margin (eg Figures 1 View Figure 1 (g), 5(g), 8(g), 10(g), 22(g)) ........................ 7
7. IOS at most 0.8× eye length; hyperoccipital carina medially pointed in between posterior ocelli ( Figures 1 View Figure 1 (g), 24(d)) ............................................................................................. 8
- IOS at least 1.1× eye length; hyperoccipital carina evenly arched between posterior ocelli (eg Figures 5 View Figure 5 (d), 8(d), 10(d), 22(g)) ................................................................................... 9
8. Submarginalis basally closer and distally distant from anterior margin of fore wing ( Figure 1 View Figure 1 (e)); posterior margins of T2–T4 distinctly raised ( Figure 1 View Figure 1 (a)); space between longitudinal costae on T1 rugose; T2 entirely with small shiny irregular areas between longitudinal costae; T3–T4 anteriorly rugose ( Figures 1 View Figure 1 (a), 26(a)); fore coxa brown, mid and hind coxae yellow ( Figure 1 View Figure 1 (c)). .......................... .......................... H. altmannae sp. n.
- Submarginalis distant from anterior margin of fore wing the entire length ( Figure 24 View Figure 24 (e)); posterior margins of T2–T4 not raised; space between longitudinal costae on T1 almost smooth; space between longitudinal costae on T2 with irregular depressions, posteriorly almost smooth ( Figure 24 View Figure 24 (e)); T3 – T4 anteriorly foveate ( Figures 24 View Figure 24 (a), 27 (e)); all coxae brown ( Figures 23 View Figure 23 (a), 24(c)) ................................ H. wintes (Kozlov and Lê)
9. T2 with depressions sometimes interspersed with longitudinal costae medially ( Figures 26 View Figure 26 (I,J,L,M), 27(d)) ............................................................................................................... 10
- T2 with reticulations or areolae, sometimes interspersed with longitudinal costae ( Figures 26 View Figure 26 (d,g,n,p), 27(a)) .............................................................................................................. 14
10. T2 basally with elongate depressions, at least 5× as long as wide followed by polygonal depressions, at least 2–3× as long as wide; T1 with ill-defined longitudinal costae ( Figure 26 View Figure 26 (j)) ........................................... ........................................... H. irregularis sp. n.
- T2 basally with short depressions, at most 2× as long as wide followed by depressions which are round or transverse in shape; T1 with percurrent longitudinal costae ( Figures 26 View Figure 26 (i,l,m,d)) ............................................................................................................................ 11
11. Submedian carina of frontal depression restricted to dorsal 2/5 of frontal depression ( Figures 10 View Figure 10 (f), 14(f)); T2 medially with only depressions, at most with short longitudinal costae on anterior margin ( Figure 26 View Figure 26 (i,m)); lateral propodeal area with setae posteriorly ( Figures 10 View Figure 10 (g), 14(g)) ................................................................................................. 12
- Submedian carina extending beyond half the length of dorsal frontal depression ( Figures 13 View Figure 13 (f), 22(f)); T2 with depressions interspersed with longitudinal costae medially, extending almost entire length of tergite ( Figures 26 View Figure 26 (l), 27(d)); lateral propodeal area without setae ( Figures 13 View Figure 13 (g), 22(g)) ................................................................................. 13
12. Stigmalis of fore wing incurved; hind wing wide, at most 4× as long as wide ( Figure 10 View Figure 10 (e)); depressions on T2 small and closely spaced; T1 anteriorly weakly convex ( Figure 26 View Figure 26 (I)); central keel on frons extends>4/5 length of frontal depression ( Figure 10 View Figure 10 (f)) ... ........................................................................................................................................ H. hrdyae sp. n.
- Stigmalis of fore wing straight, not incurved; hind wing narrow, at least 6.5× as long as wide ( Figure 14 View Figure 14 (E)); depressions on T2 large and widely spaced; T1 anteromedially projecting ( Figure 26 View Figure 26 (m)); central keel on frons extending <¼ length of frontal depression ( Figure 14 View Figure 14 (f)) .................................................................................. H. randhra sp. n.
13. Longitudinal costae on T2 unevenly distributed ( Figures 13 View Figure 13 (a), 26(l)); setae on metapleuron sparse ( Figure 13 View Figure 13 (c)); submarginalis distant from anterior margin of fore wing the entire distance, and without a dip before reaching marginalis ( Figure 13 View Figure 13 (e)) .......... ........................................................................................................................................ H. peludo sp. n.
- Longitudinal costae on T2 evenly distributed ( Figures 22 View Figure 22 (a), 27(d)); setae on metapleuron dense ( Figure 22 View Figure 22 (c)); submarginalis close to anterior margin of fore wing in basal 7/10, and with a slight dip before reaching marginalis ( Figure 22 View Figure 22 (e)) ..................... .................................................................................................................... H. westeberhardae sp. n.
14. T2 with large reticulations at least 0.5× MOD, all reticulations of even shape and size ( Figures 5 View Figure 5 (a), 26(d)) ............................................................................................ H. cookeae sp. n.
- T2 either areolate or with small reticulations (at most 0.2× MOD) of uneven shape and size (eg Figures 26 View Figure 26 (g,i,j,n), 27(d)) ................................................................................................. 15
15. Hind wing posteromedially projecting ( Figure 8 View Figure 8 (e)); T2 with unevenly shaped areolae ( Figures 8 View Figure 8 (a), 26(g)); metapleuron densely setose ( Figure 8 View Figure 8 (c))..... H. goodallae sp. n.
- Hind wing posteromedially evenly rounded ( Figures 15 View Figure 15 (e), 17(e), 18(e)); T2 with reticulations ( Figures 15 View Figure 15 (a), 17(a), 18(a), 26(n,p), 27(a)); metapleuron sparsely setose ( Figures 15 View Figure 15 (c), 17(c), 18(c)) ............................................................................................................. 16
16. Femoral depression entirely transversely carinate; mesepimeral sulcus not foveate ( Figure 15 View Figure 15 (c)); submarginalis distant from anterior margin of wing the entire length ( Figure 15 View Figure 15 (e)) ................................................. ................................................. H. samantara sp. n.
- Femoral depression transversely carinate interspersed with foveae; mesepimeral sulcus distinctly foveate ( Figures 17 View Figure 17 (c), 18(c)); submarginalis adjacent to anterior margin of wing in basal half ( Figures 17 View Figure 17 (e), 18(e)) .............................................................. 17
17. Central keel at least half the length of frontal depression; transverse carinae in frontal depression closely spaced ( Figure 18 View Figure 18 (f)); mesoscutum and mesoscutellum without longitudinal carinae ( Figure 18 View Figure 18 (a)); ventral mesopleuron narrow, half the width of dorsal mesopleuron ( Figure 18 View Figure 18 (c)) .................................................................. H. taraka sp. n.
- Central keel short, at most 0.1× length of frontal depression; transverse carinae in frontal depression spaced apart ( Figure 17 View Figure 17 (f)); mesoscutum and mesoscutellum with longitudinal carinae ( Figure 17 View Figure 17 (a)); ventral mesopleuron wide,>0.8× the width of dorsal mesopleuron ( Figure 17 View Figure 17 (c)) .................................................................... H. stupa sp. n.
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