Fortipalpa heredia, Supeleto & Santos & Aguiar, 2022

Supeleto, Fernanda A., Santos, Bernardo F. & Aguiar, Alexandre P., 2022, Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae), Zootaxa 5219 (6), pp. 501-533 : 512-515

Fortipalpa heredia sp. nov.

( Figs 6–7 , 19–21 )

Description. Holotype ♀. Fore wing 6.90 mm. Head ( Figs 6 , 7A–B ). MLW 1.71. MWW 0.50. MSM 0.33. CHW 0.54. Clypeus anterior margin centrally with two small tubercles. Antenna with 22 flagellomeres. Supra-antennal area polished smooth; median line distinct and straight.

Thorax ( Figs 6 , 7A ). Epomia approximately straight. Mesoscutum length to width 0.96 × as long as wide. Notaulus reaching about 0.51 of mesoscutum length. Scuto-scutellar groove with vestigial longitudinal wrinkles. Epicnemial carina reaching about 0.6 of distance to subalar ridge; ending at pronotal spiracle; distinctly curved. Sternaulus moderately strong, wide and shallow at posterior 0.25; weakly sinuous. Mesopleuron entirely polished smooth. Mesosternum medio-longitudinal suture without transverse wrinkles. Transverse furrow anterior to propodeum entirely polished smooth. Metapleuron densely punctate; glabrate.

Propodeum ( Figs 6 , 7A&C ).Area behind anterior transverse carina transversally strigate, arched posteriorwards. Spiracle elliptic, SLW 1.88. Anterior transverse carina medially slightly arched forwards.

Wings ( Fig. 6 ). Fore wing crossvein 1m-cu more or less uniformly curved; limit between 1m-cu and 1-Rs+M more or less traceable. Vein 2Cua 0.64 × as long as crossvein 2cu-a. APH 0.86. Areolet subpentagonal; distinctly wider than high, AWH 0.62. Crossvein 3r-m spectral. Crossvein 2r-m slightly shorter than 3r-m. Vein 3-M distinctly shorter than 2-M. HW1C 1.07. Hind wing vein Cub straight. Vein 2-1A ending at 0.18 of distance to wing margin.

Metasoma ( Figs 6 , 7A&D ). T1LW 2.51. T1WW 2.07. T1 spiracle at 0.56 of its length. Median depression of T1 subcircular, distinct. T2LW 0.97. T2WW 2.20. T2–8 faintly coriarious. Ovipositor straight, OST 1.53; ventral valve apex with 14 teeth ( Fig. 7D ).

Color. Head whitish (249,242,225) with supra-antennal area and vertex black (36,31,21). Clypeus mid ventrally and dupraclypeal area latero ventrally with brown spot. Malar space black. Orbital band almost complete, briefly interrupted near 6 h. Mandible basally whitish, apical 0.65 dark brown. Antenna dark brown (62,39,22) with whitish band composed by fl5 apical 0.70 whitish, fl6–13, and fl13 dorsally. Pronotum black, dorso anterior spot and collar whitish. Propleuron whitish. Mesoscutum black, median lobe centrally with whitish spot. Tegula and subalar ridge whitish. Epicnemium dorsally dark brown, ventrally yellow (245,197,66). Mesopleuron yellow, mid anteriorly with whitish (241,229,195) area, postero ventrally with whitish spot, dorso anteriorly with large dark brown area; hypoepimeron whitish. Mesepimeron and metapleuron yellow. Dorsal division of metapleuron whitish. Scutellum and postscutellum whitish (249,242,225). Propodeum shinning brown, posteriorly with large pale yellow (233,221,143) area. Legs bright yellow (255,210,87). Fore leg: coxa yellow (245,197,66), basally with brown (124,77,49) area; trochanter, trochantellus, femur, tibia and t1 bright yellow; t2 bright yellow, apical 0.45 brown (124,77,49); t3–5 dark brown. Mid leg: coxa yellow (245,197,66); trochanter, trochantellus, femur and tibia bright yellow; t1 bright yellow, apical 0.45 brown (124,77,49); t2 brown (124,77,49); t3–5 dark brown. Hind leg: coxa yellow (245,197,66); trochanter, trochantellus, femur and tibia bright yellow; t1 bright yellow, apical 0.75 whitish; t2–4 whitish; t5 whitish, apically dark brown. Metasoma striped black and whitish (241,229,195). T1 basal 0.2 and apical 0.25 whitish, centrally dark brown (45,29,16). T2–8 bicolored, dark brown with posterior 0.35–0.50 whitish. S2–3 laterally bicolored, anterior 0.75 dark brown (62,39,22), posteriorly whitish; S4 anterior 0.45 dark brown, posteriorly whitish.

Morphometry. See Table 1, Figs 20–21 , and Appendix.

Male. Unknown.

Comments. Structure. Most similar to F. panamensis , but differs by having the following features: Mesopleuron entirely polished smooth (vs. polished smooth, centro-anteriorly with transversal striation, ventral half punctate); epicnemial carina reaching about 0.6 of distance to subalar ridge (vs. reaching about 0.8 of distance to subalar ridge); epomia short,ending far from dorsal margin of pronotum(vs. moderately long, ending near dorsal margin of pronotum); epicnemial carina distinctly curved (vs. almost straight); HW1C 1.07 (vs. 1.30–1.49); epomia after diverging from pronotal collar approximately straight (vs. distinctly curved); fore wing vein 2Cua 0.64 × as long as crossvein 2cu-a (vs. 0.71–0.83); APH 0.86 (vs. 0.47–0.76); epicnemial carina ending at pronotal spiracle (vs. ending above pronotoal spiracle); metapleuron glabrate (vs. moderately pilose); T1LW 2.51 (vs. 2.67–2.85); median depression of T1 subcircular, distinct (vs. elongate, very shallow); T2–8 faintly coriarious (vs. from distinctly coriarious T2 to smooth T8); hind wing vein Cub straight(vs. distinctly convex on posterior half, otherwise straight);AWH0.62 (vs. 0.71–0.96).

Also similar to F. frida , but differs by having the following features: clypeus anterior margin centrally with two small tubercles (vs. without teeth); epomia short, ending far from dorsal margin of pronotum (vs. moderately long, ending near dorsal margin of pronotum); epicnemial carina reaching about 0.6 of distance to subalar ridge (vs. reaching about 0.7 of distance to subalar ridge); MLW 1.71 (vs. 1.45–1.53); HW1C 1.07 (vs. 1.70–2.00); APH 0.86 (vs. 0.69); sternaulus weakly sinuous (vs. strongly sinuous); mesopleuron entirely polished smooth (vs. polished smooth, ventral half punctate); epomia after diverging from pronotal collar approximately straight (vs. distinctly curved); metapleuron glabrate (vs. moderately pilose); limit between fore wing crossvein1m-cu and 1-Rs+M more or less traceable (vs. fully continuous with 1-Rs+M); median depression of T1 subcircular, distinct (vs. elongate, very shallow); T2–8 faintly coriarious (vs. from distinctly coriarious T2 to smooth T8); hind wing vein Cub straight (vs. almost straight, very slightly convex); OST 1.53 (vs. 1.25–1.38); AWH 0.62 (vs. 0.71–0.75); supra-antennal median line distinct and straight (vs. with slight median line).

Also similar to F. exelysae , from which it differs by having the following features: Epicnemial carina reaching about 0.6 of distance to subalar ridge (vs. reaching about 0.8 of distance to subalar ridge); APH 0.86 (vs. 0.60– 0.70); area behind anterior transverse carina transversally strigate, arched posteriorwards (vs. transversally strigate, generally weak, antero-laterally faint).mesopleuron entirely polished smooth (vs. polished smooth, centro-anteriorly with transversal striation, ventral half punctate); median depression of T1 subcircular, distinct (vs. drop-shaped, distinct, large); hind wing vein Cub straight (vs. almost straight, very slightly convex); scuto-scutellar groove with vestigial longitudinal wrinkles (vs. with distinct longitudinal wrinkles); metapleuron glabrate (vs. moderately pilose); supra-antennal area polished smooth (vs. distinctly coriarious).

Color pattern. Closest to F. yucatanica , from which it differs by having hind coxa dorsally without whitish spot (vs. dorsally with distinct whitish spot); propodeum brown (vs. orange); clypeus mid ventrally with dark brown spot (vs. whitish); supraclypeal area latero ventrally with small brown spot (vs. whitish); scape ventrally dark brown (vs. whitish); and ovipositor yellowish (vs. brown).

Also similar to F. exelysae , from which it differs by having ovipositor yellowish (vs. brown); propodeum brown (vs. orange); and dorsal margin of pronotum anteriorly with whitish spot (vs. entirely yellow).

Etymology. The specific epithet is a literal reference to the municipality where the holotype was collected. Distribution records. Costa Rica ( Fig. 19 ).

Material examined. One specimen, 1♀. Holotype: COSTA RICA, 1♀, Heredia, Santa Rosa, 2?, 1.100 m, 9°58′10″N, 84°05′54″W, 1981-8, M.Fischel,?, FAS 4628 ,?, TAMU.

Figures

FIGURE 6. Fortipalpa heredia sp. nov. Holotype female. Habitus.

FIGURE 7. Fortipalpa heredia sp. nov. Holotype female. A, Dorsal view. B, Head, frontal. C, Propodeum, dorsal. D, Ovipositor tip, left side.

FIGURE 19. All known distribution records for Fortipalpa spp. (first map) and individual records for each species. Taxa presented in alphabetical order.

FIGURE 20. All results for morphometric ratios calculated for specimens of Fortipalpa spp., shown comparatively. On each plot, the species are sorted on the Y-axis from highest (top) to lowest (bottom) average. Graphs organized by alphabetical order of ratios.

FIGURE 21. Continuation of Figure 20.