Mannerheimia emodensa Coiffait, 1977

Shavrin, Alexey V., 2024, On some Himalayan species of the genus Mannerheimia Mäklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini), Zootaxa 5474 (5), pp. 584-592 : 587-589

Mannerheimia emodensa Coiffait, 1977

( Figs 3 , 9–10 )

Mannerheimia emodensa Coiffait, 1977: 266 ; Coiffait 1982: 150, Shavrin 2021: 307

Type material examined. Holotype ♂ ( Fig. 3 ; dissected prior to the present study: the aedeagus and the apical genital segment are glued on the same card with the specimen): ‘Umg.Alm Darghari | b. Maharigaon, 4000m’ <printed>, ‘Gebiet von Jumla | Westnepal,lg.H.Franz’ <printed>, ‘TYPE’ <red, printed>, ‘ Mannerheimia | emodensa | H. COIFFAIT det. 19[printed]76’ <handwritten in black> ( NMW). Paratypes: 3 ♂♂ (one specimen dissected), 2 ♀♀: first two labels as that in the holotype, with additional red printed label: ‘PARATYPE’ ( NMW). All types with additional printed label: ‘ Mannerheimia | emodensa Coiffait, 1977 | Shavrin A. V. det. 2024’.

Additional material examined. NEPAL: 9 ♂♂, 3 ♀♀: same data as the holotype ( NMW) ; 1 ♂: ‘ Dzunda Khola-Tal b. Talphi 3000-3500m’, ‘ Gebiet von Jumla Westnepal,lg. H.Franz’ ( NMW) ; 1 ♀: ‘ Dampelek b. Jumla’, ‘ Gebiet des Rarasees Westnepal , lg. H. Franz’ ( NMW) .

Redescription. Measurements (n=20): HW: 0.47–0.53; HL: 0.31–0.33; OL: 0.14–0.15; AL (holotype): 0.85; PL: 0.33–0.37; PW: 0.61–0.68; ESL: 0.74–0.78; EW: 0.90–1.05; MTbL (holotype): 0.52; MTrL (holotype): 0.30 (MTrL 1–4: 0.17; MTrL 5: 0.13); AW: 0.95–1.03; AedL: 0.36–0.40; BL: 1.98–2.65 (holotype: 2.37).

Habitus as in Fig. 3 . Body yellow-brown to reddish-brown, with distinctly darker head (abdomen of some specimens slightly darker than forebody; lateral and basal portions of pronotum and apical portion of elytra slightly paler in holotype and in some paratypes); antennomeres 5–11 or 6–11 brown; mouthparts, antennomeres 1–4 or 1–5 and paratergites yellow. Forebody without microsculpture; abdomen with fine and dense transverse microreticulation. Head with moderately fine and sparse punctation, distinctly denser in middle; punctation of pronotum denser, larger and deeper than that in middle portion of head, finer and sparser in middle, and sometimes with narrow impunctate portion in middle or in mediobasal third; scutellum with one to four fine punctures in middle or without them; punctation of elytra denser, slightly larger and deeper than that on pronotum, punctures in middle of each elytron forming five to six vague and tangled rows of serial punctures, but some specimens with markedly coarse surface without longitudinal rows between punctures.

Head 1.5–1.6 times as broad as long, slightly elevated in middle, with very short and sometimes indistinct anteocellar foveae as long as diameter of ocellus. Antennomere 3 narrower and slightly shorter than 2, 4 distinctly shorter than 3 and slightly longer than broad, 5–6 indistinctly broader than 4, 7–8 slightly longer and distinctly broader than 6, 9–10 slightly longer and broader than 8, apical antennomere about as long as two preceding antennomeres or slightly shorter.

Pronotum 1.8 times as broad as long, 1.2 times as broad as head, widest slightly above middle, evenly narrowed toward obtuse posterior angles, sometimes with indistinctly sinuate laterobasal margins; lateral portions slightly impressed, distinctly broader and deeper in laterobasal portions.

Elytra 1.2–1.3 times broader than long, slightly or strongly widened apicad.

Metatarsus 1.7 times as long as metatibia.

Abdomen about as broad as elytra, with two small oval or transverse tomentose wing-folding patches in middle of abdominal tergite V.

Male. Apical margin of abdominal tergite VIII straight or slightly sinuate. Apical margin of abdominal sternite VIII widely sinuate. Aedeagus with wide basal portion, gradually narrowed apically, with relatively narrow median lobe and small rounded apex; parameres with wide apical portions, slightly exceeding apex of median lobe; internal sac narrow and moderately long ( Fig. 9 ). Lateral aspect of aedeagus as in Fig. 10 .

Female. Apical margin of abdominal tergite VIII straight or somewhat rounded. Apical margin of abdominal sternite VIII rounded.

Comparative notes. Based on the general shape of the body and the aedeagus, M. emodensa is similar to M. cachemirica , from which can be distinguished by the slightly broader elytra, larger and denser punctation on the pronotum and elytra, slightly shorter preapical antennomeres, and longer parameres with significantly broader apical portions.

Distribution. Mannerheimia emodensa is known from four localities in western Nepal.

Bionomics. Specimens were collected at altitudes from 3000 to 4000 m a.s.l. Detailed bionomical data are unknown.

Remarks. Mannerheimia emodensa based on several specimens was originally described from four localities in “..région de Jumla”, western Nepal. I studied the holotype and five paratypes from NMW, and several specimens without paratype label, some of which are apparently belonging to the type serie (see Additional material section above). Coiffait (1982) included it to the key of Mannerheimia of the Himalayan Region. Shavrin (2021) included this species to the key of Mannerheimia .

Figures

FIGURES 1–5. Habitus of Mannerheimia (holotypes): 1–M. cachemirica, 2–M. vicina, 3–M. emodensa, 4–M. obscura, 5–M. scutellata, Scale bar: 1.0 mm.

FIGURES 6–10. Aedeagus of Mannerheimia (Figs 6–7: M. cachemirica (holotype); Fig. 8: M. vicina (holotype); Figs. 9–10: M. emodensa (Nepal: Maharigaon): 6, 8–9–parameral view, 7, 10–lateral view. Scale bars: 0.1 mm.