Drymeia Meigen, 1826

Pont, Adrian C., 2022, The genus Drymeia Meigen in the Caucasus Mountains, with a note on the identity of Aspilia glacialis Rondani, 1866 (Diptera: Muscidae), Zootaxa 5134 (3), pp. 355-382 : 356-357

Key to Caucasus species of Drymeia Meigen

1 Facial edge not or only a little projecting forward beyond the level of the profrons when the head is viewed in profile ( Fig. 22 ). Scutellum on the lateral margins, below the level of the strong setae, with 5–10 setulae in 1–2 rows. ♂: prealar seta absent or short, shorter than 2nd notopleural seta (present in some D. vicana ). ♀: hind tibia without a posteroventral apical seta..... 2

- Facial edge projecting forward far beyond the level of the profrons when the head is viewed in profile ( Fig. 12 ). Scutellum bare on lateral margins, but with a row of setulae on the ventral surface in D. sororcula . ♂: prealar seta present and strong. ♀: hind tibia with a posteroventral apical seta..................................................................... 4

2 Anepimeron setulose. Katepimeron with a few short setulae. (♂ with the tip of the abdomen forming a “basket”, with numerous dense setae on the apical margin of tergite 5 and on subsequent tergites, enclosing the genital capsule; mid femur without posteroventral setae except for a group of fine curled setae at base, some longer than femoral depth; hind femur with a row of anteroventral setae in apical half but without posteroventrals except at base).................... D. caucasica (Schnabl)

- Anepimeron bare. Katepimeron bare...................................................................... 3

3 Eyes densely haired, less conspicuously so in female. ♂: fore leg with no posteroventral setulae at apex of tarsomere 1; mid femur with complete rows of fine anteroventral and posteroventral setae, the posteroventrals longer; hind femur with complete rows of anteroventral and posteroventral setae................................................. D. vicana (Harris)

- Eyes bare. ♂: fore tarsomere 1 with several long posteroventral setulae near apex; mid femur without posteroventral setae, and without anteroventrals except for some long, bent setae on basal quarter and some long anterior setae above them ( Figs 4–5 ); hind femur with anteroventral setae in apical half but without posteroventrals..................... D. setibasis (Huckett)

4 Parafacial very broad, at middle almost twice as wide as width of postpedicel ( Fig. 12 ). Face with a prominent snout between the base of the antennae. (3 postsutural dorsocentrals.) ♂: hind tibia with a posteroventral apical seta, but with no prolongation of the apex of the tibia to form a short spur ( Fig. 13 ). (♂ with complete rows of anteroventral and posteroventral setae on mid femur and hind femur, these rows also complete but sparser in the ♀)............................ D. fratercula sp.nov.

- Parafacial narrower, at middle equal to width of postpedicel ( Fig. 33 ). Face without a snout between the antennae. ♂: hind tibia without a posteroventral apical seta, but with a ventral spur-like prolongation of the apex of the tibia................... 5

5 4 postsutural dorsocentral setae. Scutellum bare on ventral surface.Anepimeron always bare. ♂: mid femur with the anteroventral setae restricted to a group of 5–6 setae in apical fifth, and with posteroventral setae in basal two-thirds ( Fig. 5 ); hind femur with a complete row of anteroventrals, but with no posteroventrals. ♀: parafacial with a shining patch at the level of antennal insertion; mid femur with a few posteroventral setae in basal half, and with 2–3 anteroventrals before apex and, sometimes, a further seta near base of femur ( Fig. 6 ); hind femur with a complete anteroventral row as in ♂ ....... D. fasciculata (Stein)

- 3 postsutural dorsocentral setae. Scutellum with several strong setulae on ventral surface.Anepimeron often with a few setulae. ♂: mid and hind femora each with complete rows of anteroventral and posteroventral setae. ♀: parafacial without a shining patch in upper part; mid femur without posteroventral setae, on anteroventral surface with 1 seta near base and 2 very short, almost spinulose setae just before apex; hind femur with a sparse row of anteroventrals.............. D. sororcula sp.nov.

Figures

FIGURES 22–25. Drymeia setibasis (Huckett). 22. ♂ head, lateral view. 23. ♂ mid femur, anterior view. 24. ♀ mid femur, an- terior view. 25. ♂ hind tibia and tarsomere 1, anterior view. Scale-line 1 mm (3) and 0.5 mm (4–6). (Georgia, Kazbegi, Gergeti, scree & river, 2950–3200 m, 11.vii.1983.)

FIGURES 12–16. Drymeia fratercula sp.nov. 12. ♂ head, lateral view. 13. ♂ hind tibia and tarsomere 1, posterior view. 14. ♀ fore tarsomeres, anterior view. 15. ♂ sternite 5, dorsal view. 16. ♀ tergite 8. Scale-line 0.25 mm (15), 0.5 mm (16). (Georgia, Ka- zbegi, Mt Koltesh, meadow/heath, 2300 m, 8.vii.1983 (12, 13) and 30.vi.1983 (14, 16); Mt Koltesh, heath, 2500 m, 30.vi.1983 (15).)

FIGURES 3–4. Drymeia fasciculata (Stein), ♂ habitus. 3. Lateral view. 4. Dorsal view. (Georgia, Kazbegi, Mt Sameba, pasture, 2250–2300 m, 2.vii.1983.)

FIGURES 5–7. Drymeia fasciculata (Stein). 5. ♂ mid femur, anterior view. 6. ♀ mid femur, anterior view. 7. ♂ sternite 5, dorsal view. Scale line 0.25 mm. (Azerbaijan.)

FIGURES 33–35. Drymeia sororcula sp. nov. 33. ♂ head, lateral view. 34. ♀ fore tarsomeres, posterior view. 35. ♂ sternite 5, dorsal view. Scale line 0.25 mm. (Georgia, Kazbegi, Mt Kuro, heath, 2120–2330 m, 29.vi.1983 (33, 35), Mt Koltesh, meadows, 2000–2100 m, 8.vii.1983 (34).)