Thysanomeros jacobyi (Lefèvre), Lefevre

Flowers, R. Wills, 2003, A new genus and a new Costa Rican species of Endocephalites (Chrysomelidae: Eumolpinae: Eumolpini), Zootaxa 221, pp. 1-11 : 5-7

Thysanomeros jacobyi (Lefèvre)

( Figs. 2, 4, 6, 7 , 8, 9 , 13, 14 , 18 )

Prionodera jacobyi Lefèvre 1885 :cxc, new combination

Male. Body dorsally convex; length 5.6 – 6.5 mm. Head and pronotum yellowish brown, elytra metallic green or bluish green; antenna with scape dark brown, washed with black, antennomeres 3­11 black. Underside yellowish brown with femora yellowish brown, tibia brown in basal fourth, dark brown to black in remainder, tarsi dark brown to black. Head with apical margin of clypeus deeply emarginate, antennal calli swollen. Mouthparts dark reddish brown, apex of labrum emarginate with 4 dorsal setae. Prothorax distinctly wider than long, L/W = 0.72; lateral margin narrow undulate, forming one to three teeth at the mid­point ( Fig. 4 ). Greatest width of pronotum at middle; disc with scattered fine punctures at middle. Undersurface of thorax smooth. Prosternum longitudinally strongly inclined behind coxae, depressed in middle at posterior margin; posterior margin of intercoxal process very shallowly concave, width of intercoxal process 0.6 x diameter of procoxa. Pro­ and mesofemora with erect setae on ventral margin in basal fourth, metafemur with dense brush of ventral setae in basal half ( Fig. 2 ). Scutellum V­shaped: Abdomen with all segments covered with erect yellow setae. Male sternum VII with a weak depression in center. Pygidial surface smooth, covered with long, erect setae. Median lobe of aedeagus in lateral view smoothly curved ( Fig. 8 ); apex with a pointed protuberance thickened ventrally; basal hood long, lightly sclerotized, with apodemes indistinct at lateral margins of hood. Apex of median lobe in apical view subtruncate, minutely acuminate ( Fig. 9 ). Endophallus membranous, moderately elongate, thicker and lobed dorsally; apical sclerite of endophallus straight, with apical and basal curved flaps ( Fig. 8 ).

Female. Body oval; length 6.5 – 8.3 mm; head, and pronotum as in male, elytra metallic green or dark blue, underside and antennae as in male, tibiae ranging from entirely yellowish brown to entirely black. Head with labrum, frons, clypeus, eyes and antennae similar to male; mouthparts similar to male. Prothorax distinctly wider than long, L/W = 0.62, intercoxal process 0.7 x diameter of procoxa. Legs similar to male but fewer setae on ventral margins of femora, metafemur lacking dense setal brush. Scutellum U­shaped: Abdominal sterna covered with erect setae, a slightly larger seta on each side of midline. Sternum VII with numerous long lateral and apical setae. Abdominal segments VIII­XI ( Fig. 13 ) forming short ovipositor. Baculum distinct, apical, subequal to gonocoxae. Gonocoxae short, robust, with long setae in apical half, a membraneous area in basal half; coxostyli very small, with several long apical setae. Spermatheca as in Fig. 14 ; receptacle tapered in basal fourth, spermathecal duct thin, sclerotized, loosely coiled.

Specimens Examined: (41 ɗ; 35 Ψ) COSTA RICA, Alajuela Prov: 1 Ψ Río San Lorencito, 900m Res. For. San Ramón, 5 km N Col. Palmarea, Mar 1990, Curso Carabidae , L­ N 244500_470700; Guanacaste Prov: 1ɗ, Maritza, 600 m, W side Volcán Orosi. Malaise Tp 1989. GNP Biod. Sur, L_N_326900_373000 #6658; 1 Ψ, Est. Maritza, 600m, Lado oeste del Volcn Orosi, II curso Parataxonomos, Ago 1990, L­ N 326900_373000; Puntarenas Prov.: 13 ɗ, 6 Ψ; Estac. Quebrada Bonita, 50m, R. B. Carara, May 1990. R. Zuiga. L­ N 194500_469850; 1ɗ, same locality and collector, Jun 1990; 2 ɗ, same locality and collector, Jun 1991; 1 Ψ, same locality, Jun. 1993, R. Guzman; 1 Ψ, Corcovado National Park Osa Peninsula. 5­9 May 1978 D.H. Janzen; 1 Ψ, Est. Sirena, 0 ­ 100 m, P. N. Corcovado, 1991. Tp Malaise, L S 270500,508300; 2 Ψ, same locality, J. C. Saborio, Jun 1991; 1 Ψ, same locality, Tp Malaise Mar ­ Jun 1991; 3 ɗ, 6 Ψ, same locality, G. Fonseca, May 1992; 1 Ψ, same locality and collector, Jul 1992; 11 ɗ, 5 Ψ, same locality and collector, May 1993; 2 ɗ, 1 Ψ, same locality and collector, Jun 1993; 7 ɗ, 4 Ψ, same locality and collector, May 1994; 1ɗ, Est. Sirena, ACOSA, 1­ 100m. 6­12 ABR 1995. A. Picado, L N 270500 507900 #4573; 2 Ψ, Est. Sirena, Sendero Ollas (ACOSA), 150m. 20 ABR 1995. B. Gamboa, L S 272100 509400 #4744; 1 Ψ, Fila Matahambre, Fca. La Purruja, Cerro La Torre, Pen. Osa, 200 m. 4­9 May 1994, M. Segura, L N 277000_527000 # 2936; 1 Ψ, Rancho Quemado, Pen. de Osa, Prov. Punta, COSTA RICA. 200m. 1­27 Ene 1992, A. Marin, L S 292500_511000 #1779.

Figures

FIGURE 18. Distribution of Thysanomeros. T. jacobyi, closed circles. T. ulateae, closed squares.

FIGURES 2 ­ 7. External characters of Thysanomeros. 2 – 3. Hind femur of male. 2. T. jacobyi. 3. T. ulateae. 4 – 5. Lateral margin of pronotum. 4. T. jacobyi, 5. T. ulateae. 6. Fore tarsi of T. jacobyi: left, male; right, female. 7. Pygidium of T. jacobyi.

FIGURES 8 ­ 12. Male genitalia of Thysanomeros. 8, 10. Median lobe. 8 T. jacobyi, showing endophalus and apical sclerite. 10. T. ulateae. 9, 12. Apex of median lobe. 9. T. jacobyi. 12. T. ulateae. 11. Apical sclerite, T. ulateae.

FIGURES 13 ­ 17. Female characters of Thysanomeros. 13, 16 ­ 17. ovipositors. 13. T. jacobyi, ventral. 16. T. ulateae, ventral. 17. T. ulateae, dorsal. 14 ­ 15. spermatheca. 14. T. jacobyi. 15. T. ulateae.