Mycomyiella tannerorum, Gammelmo, Øivind, 2004

Gammelmo, Øivind, 2004, A review of the Afrotropical genus Mycomyiella Matile, 1973 (Diptera, Mycetophilidae, Mycomyini), with the description of six new species, Zootaxa 625, pp. 1-30 : 26-27

Mycomyiella tannerorum new species

(Figs 23–26)

Type material: Holotype male, TANZANIA, Tanga region, West Usambara Mountains, Mazumbai, 20–26.xi.1990, ZMB's Tanzania Expedition, ( ZMUN). Paratypes: as for holotype except 1 male, 27.x.1990; 1 male, 31.x.1990; 1 male, 2–3.xi.1990; 1 male, 4.xi.1990; 1 male, 4–6.xi.1990; 1 male, 7–12.xi.1990; 1 male, 21–26.xi.1990; 2 males, 23.xi.1990; 5 males, 26–29.xi.1990; 3 males, 29.xi.–3.xii.1990.

Etymology. Named after Mrs. and Mr. Tanner, who initiated the establishment of the Mazumbai Forest Reserve.

Diagnostic characters. The male groups with M. diseta new species and M. elegans new species in having a forked gonostylus and gonocoxite with long lobes. It can be separated from both species by having moderately long, straight lobes of tergite 9.

Description

Male (n=18). Total length 2.98–3.71, 3.45 mm.

Coloration: Head light brown to brown. Palpomeres and other mouthparts light brown to yellow. Ocellar area dark brown. Face yellow­brown, posterior parts of head light brown. Antennae yellow­brown; scape, pedicel and base of first flagellomere yellow. Antepronotum yellow. Scutum brown to dark brown, with 3 light yellow longitudinal bands. Anepisternum and preepisternum yellow. Scutellum light brown to brown. Laterotergite light brown to brown, mediotergite brown. Coxae yellow; femura, tibiae and tarsi light brown. Tergite 1 light brown; tergite 2 yellow with posterior, light brown band; tergite 3 brown; tergite 4 yellow with narrow brown median line; tergites 5–6 brown; tergite 7 yellow. Sternites 1–4 yellow; sternites 5–6 yellow with brown, lateral patches; sternite 7 yellow­brown. Terminalia light brown to brown.

Head: Head 0.47–0.53, 0.51 mm long; 0.50–0.55, 0.51 mm wide. First flagellomere about 1.5 times as long as wide; second flagellomere as long as wide. Clypeus 0.10–0.13, 0.12 mm long; with lateral and apical setae. Ocellar area 0.10–0.13, 0.11 mm wide; placed below upper margin of compound eye. Some setae in front of ocellar area. Posterolateral portion of frons densely setose, setae reaching just below ocellar area. Face bare. Mouthparts elongated. Palpomere 4 with 2–5 setae subbasally, palpomere 3 with several setae in basal two­thirds. Palpomere ratios (2–5) as: 1.00: 3.25: 3.00: 5.25.

Thorax: Total length 0.75–0.85, 0.80 mm. Scutum 0.64–0.72, 0.68 mm long; scutellum 0.13–0.18, 0.17 mm long.

Legs: Leg measurements and ratios as in Tables 4 & 5.

Wings: Wing 3.00–3.32, 3.22 mm long; 1.09–1.13, 1.12 mm wide. Sc1 lacking. Sc ending in R1 slightly distal to middle of small cell. Sc bearing 7–12, 9 macrotrichia apically. R4/Rs­cell 1.0–2.0 times as long as wide. CuA fork slightly distal to M fork. Wing measurements are given in Table 6 .

Abdomen: Abdomen 2.56–2.72, 2.68 mm long; tergite 5 0.63–0.82, 0.73 mm wide.

Male terminalia (Figs 23–26). Tergite 8 wide, with several long posterolateral setae. Tergite 9 prominent, strongly setose; with long, narrow, sigmoid lobes, with setae medially. Gonocoxite with few lateral setae, or entirely bare; lobes long, with 2–4 setae; lateral prolongation large, club­shaped, setose. Gonostylus furcated in apical half, with 2–5 setae. Aedeagus prominent, covered with short setae and with long setae subapically. Sternite 8 subtriangular basally, apical one­half narrowly digitate, setose.

Female (n = 19). Total length 2.87–3.88, 3.59 mm. As for males, except:

Coloration: Tergite 2 yellow; tergite 4 brown; tergite 7 yellow­brown. Sternites 4–6 brown; sternite 7 yellow.

Head: Palpomere ratios (2–5) as: 1.00: 3.15: 3.15: 5.37.

Thorax: Total length 0.78–0.93, 0.86 mm. Scutum 0.63–0.83, 0.77 mm long; scutellum 0.14–0.15, 0.14 mm long.

Wing: Wing 2.88–3.41, 3.21 mm long; 0.99–1.21, 1.10 mm wide.

Abdomen: Abdomen 2.47–2.69, 2.49 mm long.

Female terminalia: Cercus 2 normal, pointed apically. Tergite 9 strongly pointed posterolaterally. Gonocoxite 9 prominent. Sternite 10 reduced.

Distribution: Known from the type locality in the West Usambara Mountains in North­East Tanzania only.