Mycomyiella Matile

Gammelmo, Øivind, 2004, A review of the Afrotropical genus Mycomyiella Matile, 1973 (Diptera, Mycetophilidae, Mycomyini), with the description of six new species, Zootaxa 625, pp. 1-30 : 12-14

Mycomyiella Matile

Mycomyiella Matile, 1973: 189 –213.

Type species: M. camerounensis Matile, 1973 , by original designation.

Other included species: M. diseta new species, M. elegans new species, M. ghanaensis new species, M. irwini Väisänen, 1994 , M. kaputuensis new species, M. securiculata Matile, 1973 , M. tannerorum new species, and M. wangi new species.

Diagnostic characters: The imagines can be separated from other mycetophilid genera by the combination of the following characters: i) costa ending distal to apex of R5; ii) wings without distinct markings; iii) Sc incomplete, ending in R; iv) laterotergite and mediotergite bare; v) scutellum with long setae; and vi) tibia with long setae.

Generic diagnosis

Medium sized species, total length 1.9–3.9 mm.

Coloration: Vertex generally light brown to brown. Frons, face, clypeus and mouthparts yellow to brown. Antennae yellow to brown, scape, pedicel and base of first flagellomere generally paler than rest of antennae. Scutum brown to dark brown with 3 yellow longitudinal lines medially. Scutellum brown to dark brown. Laterotergite and mediotergite yellow to brown. Coxae yellow, femura light brown to brown, tibiae and tarsi brown. Abdominal tergites and sternites variable. Terminalia yellow to brown.

Head: Subcircular in frontal view. Vertex covered with setae. Compound eyes moderately setose. Two ocelli, median ocellus lacking. Ocellar area placed above or below upper margin of compound eye. Antennae with 14 flagellomeres, inserted at middle of head, approximately of same length in males and females. Scape and pedicel moderately setose, flagellomeres densely setose. Flagellomeres longer than wide to approximately as long as wide. Frons usually with some setae in front of ocellar triangle. Frontal tubercle pointed. Face appears completely fused with clypeus or weak border discernible. Clypeus with lateral setae, occasionally also with anterior setae. Palpomere 1 reduced or fused with palpomere 2. Palpomere 3 with sensory pit subapically. Palpomere ratio (2–5) about 1.0: 3.3: 3.3: 5.7. Mouthparts elongated or reduced. Cardo present.

Thorax: Scutellum with 2 large bristles projecting posteriorly. Laterotergite and mediotergite bare. Antepronotum covered with setae. Scutum densely covered with setae, dorsomedian lines almost bare.

Wing: Membrane slightly darkened around R4/Rs­cell. R4 and A1 present. Sc1 absent. CuP weak or appears absent. CuA and M separated from wing base, branched and bellshaped. Forking point of CuA about at level with small cell. Forking point of M approximately at basal one­third of wing. C, R1, R4, R5, Sc, Rs, tb, and ta with single row of setae. In females occasionally some setae proximally on M1 and M2. C produced beyond R5. Membrane without macrotrichia.

Legs: Fore coxae covered with setae proximally and laterally at apex. Mid coxae covered with setae proximally. Hind coxae covered with setae proximally and laterally at apex. Femura densely setose. Tibia with fine, long setae arranged in rows. Tibial spurs 1:2:2. Tarsi with setae arranged in rows. Tarsus 5 with tarsal claw and reduced empodium.

Abdomen: All abdominal tergites and sternites covered with setae, except anterior part of tergite 1, tergite 7, sternite 1 and anterior part of sternite 2.

Male terminalia: Tergite 8 wide, with long posterolateral setae. Tergite 9 well developed; moderately to densely setose posteriorly; with or without long posterior lobes. Gonocoxite sparsely to densely setose laterally; with long, narrow lobes; and club­shaped or subtriangular, densely setose lateral prolongation. Gonostylus simple or forked, with or without setae. Aedeagus heavily sclerotized. Sternite 8 very large, pointed or evenly rounded posteriorly, with long posterior setae. Parameres present. Ejaculatory apodeme Yshaped.

Female terminalia: Tergites 8 and 9 with posterior setae. Tergite 10 with setae laterally and medially. Cerci two­segmented, first segment 2.0–3.0 times the length of second segment; apex of second segment pointed or evenly rounded. Hypoproct apparently bare. Sternite 10 elongated posteriorly, bare or with few weak setae. Sternite 9 apparently completely reduced. Spermathecal ducts long, slender. Gonocoxite 8 pointed, setose. Gonocoxite 9 with few strong setae.