Geodromicus bodemeyeri, Bernhauer, 1902

Shavrin, Alexey V., 2022, The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae), Zootaxa 5213 (5), pp. 451-496 : 453

Redefinition of the bodemeyeri group

Diagnosis. Body medium-sized (4.50–7.20 mm); pronotum moderately narrow, usually sharply narrowed posteriad toward subparallel-sided mediobasal portions; elytra moderately short, slightly broader than long, slightly or significantly broadened posteriad; aedeagus long, with moderately small basal part, gradually narrowed toward subacute or rounded apex, with a pair of sclerotized dorsal lobes in apical portion, without ( G. (s.str.) zwickianus ) or with two narrow and elongate middle structures, and with moderately short flagellum.

Species included. G. (s.str.) bodemeyeri Bernhauer, 1902 , G. (s.str.) championi Cameron, 1924 , G. (s.str.) zwickianus Fagel, 1976 .

Distribution. Species of the bodemeyeri group are distributed in the Palaearctic Region, and known from Causasus, Turkey, Iran and the Himalayan Region.

Bionomics. The species of the bodemeyeri group inhabit banks of rivers, streams and lakes, and can be found in gravel, under stones, etc. Specimens were collected at elevations from 180 to 3800 m a.s.l. from April to October.

Remarks. This group included G. (s.str.) zwickianus and G. (s.str.) bodemeyeri , which were included by Bordoni (1984) to major and rousi groups respectively. Based on the shape of the parameres with wide apical portions and weakly sclerotized internal sac and narrow flagellum, the bodemeyeri group is similar to the convexicollis group of the nominal subgenus Geodromicus , but differs by the presence of paired sclerotized structures in apical and middle portions (except G. (s.str.) zwickianus ) of the median lobe.

Key to bodemeyeri group

1 Pronotum 1/1 broader than long. Median lobe with long paired sclerotized structures ( Figs. 7–12 ). Body length: 5.20–7.00. Habitus as in Fig. 1 . Caucasus, Turkey......................................................... G. bodemeyeri

- Pronotum 1/2–1/5 broader than long. Median lobe with or without parallel structures............................... 2

2 Elytra longer, 1/6–1/8 longer than pronotum. Median lobe wide, with long paired sclerotized structures in middle, apical portion sharply narrowed toward apex ( Fig. 13, 15 ). Body length: 5.80–7.20. Habitus as in Fig. 2 . Pakistan, India, Nepal................................................................................................... G. championi

- Elytra shorter, 1/4–1/5 longer than promotum. Median lobe moderately narrow, without sclerotized structures in middle, apical portion gradually narrowed toward apex. Body length: 4.50–6.90. Habitus as in Figs. 3–6 . Turkey, Iran...... G. zwickianus

Figures

FIGURES 7–12. Aedeagus of Geodromicus bodemeyeri (Figs. 7–8: holotype; Fig. 9: Turkey, Kagizman; Fig. 10: Turkey, Bolghar Dagh; Fig. 11: Russia, Naltshik; Fig. 12: Russia, Alagir): 7, 9–12—parameral view, 8—lateral view. Scale bars: 0.1 mm.

FIGURES 1–6. Habitus of Geodromicus: 1—G. bodemeyeri (Turkey, Kagizman), 2—G. championi (lectotype), 3—G. iranicus (holotype), 4–6—G. zwickianus (Fig. 4: Iran, Qohrud Mts.; Fig. 5: Iran, Küh-e Hezâr; Fig. 6: Turkey, Elmali). Scale bars: 1.0 mm.

FIGURES 13–18. Aedeagus of Geodromicus championi (Figs. 13–14: Nepal, Karnali; Fig. 15—Pakistan, Chitral Bumburet) and G. zwickianus (Figs. 16–17: Turkey, Yüksekova; Fig. 18: Turkey, Elmali): 13, 15–16, 18—parameral view, 14, 17—lateral view. Scale bars: 0.1 mm.