Banyallarga Navás

Prather Table Of Contents, Aysha L., 2004, Revision of the Neotropical caddisfly genus Banyallarga (Trichoptera: Calamoceratidae), Zootaxa 435 (1), pp. 1-76 : 10-11

Genus Banyallarga Navás

Banyallarga Navás 1916:78 [Type species: Banyallarga testacea Navás 1916 , original designation]. — Fischer 1963:175 [in Hydropsychidae : Macronematinae ]. — Botosaneanu & Flint 1982:24 [larva, as Phylloicus ]. — Flint 1983:77 [to Calamoceratidae ]. — Flint & Angrisano 1985:688 [larva, pupa, distinguished from Phylloicus ]. — Flint et al. 1999a:15 [checklist].

The type specimen of B. testacea (the type species of the genus) is lost, and the original description and illustration are too general for species discrimination. Therefore, in order to objectively define the genus, I am designating a neotype for B. testacea . A detailed discussion of B. testacea ís identity and the neotype designation are to be found under the heading for B. vicaria .

Generic description, adult. Forewing length 8.4–20.8 mm; female usually slightly larger than male.

Forewing color (membrane and setae) variable, overall color ranging from golden brown to dark brown, with, in some species, mottled pattern formed by hairs in various shades of gold and brown. Antennae usually twice wing length or longer (in B. loxana¸ relative antennal length of females is reduced); scape shorter than head length, round. Maxillary palps 5­segmented in both sexes; all segments 3–4 times longer than wide. Labial palps 3­segmented. Head with anterior, anteromesal, posterior and posterolateral setal warts; central setal area bare or with fine setae only, these not aggregated as distinct warts; anteromesal setal warts prominent and single or paired; posteromesal ridge present at posterodorsal margin of head ( Fig. 1A ). Mesoscutum without distinct setal warts, with stout setae arranged in two anterior­posterior sublateral lines, each line one to several setae wide; in some species additional setae scattered between these lines ( Fig. 1B ). Anterior margin of mesoscutellum straight; without warts or prominent setae ( Fig. 1B ). Metathoracic leg of males often with posterior fringe of long setae. Forewing venation typical for family: forks I–V present, discoidal and medial cells closed; R 1 recurrent onto R 2 or free to wing margin; Cu 2 and A 1 free to wing margin ( Figs. 2A ). Hind wing with forks I, II, III, and V present, discoidal and medial cells open ( Figs. 2B ); base of hind wing with semimembranous sleeve or pouch, enclosing brush of long setae ( Fig. 32B ; not present in a few species).

Male. Tergum IX much shorter than sternum IX ( Fig. 3A ); posterior margin of tergum variously modified by short projections or processes. Sternum IX narrower mesally than laterally, internal ridges may be present mesally or sublaterally; patches of pleural setae may be present dorsolaterally or ventrolaterally ( Fig. 3A ). Preanal appendages of varying length and shape, bearing most setae on apical two­thirds. Tergum X approximately equal in length to inferior appendages, posterior margin of tergum X variously modified with processes and projections. Inferior appendages 2­segmented, not highly modified; coxopodite covered with long setae, particularly on lateral and ventral surfaces; harpago cylindrical or tapered apically, bearing short, pointed, peglike setae on mesal and apicoventral surfaces; some short fine setae may be present basally on harpago; base of inferior appendages occupies 1/2 to 2/3 length of sternum IX. Phallobase a simple curved tube, curvature consistent throughout genus, phallotremal sclerites reduced; dorsal sclerite U­ or Vshaped; ventral sclerite narrow and flat or absent; endotheca may consist of multiple membranous lobes with species­specific arrangement.

Female. Sternum VIII more darkly sclerotized than anterior segments, anterior margin marked by darkly sclerotized ridge; posterior margin bearing many short stout setae, mesally cleft to anterior ridge; lateral margins indistinct beyond lateral apodemes extending from anterior margin. Tergum IX with anterior margin entire and concave; mesally without clear junction with tergum X. Sternum IX with anterior and posterior pair of striate lobes lateral to vaginal opening. Tergum X bearing paired posterolateral appendages, varying in shape, length and degree of distinctness from tergite; these appendages bearing many long setae apically; posteromesal portion of tergite membranous or semisclerotized. Sternum X mostly membranous except for bases of dorsal appendages; semisclerotized patches and short fine setae may be present lateral to anal opening. Vaginal apparatus consisting of anterior and posterior sclerites; duct leading to spermatheca arises from within sclerotized pocket on ventral surface of anterior sclerite; duct leading to bursa copulatrix arises from anterior margin; posterior sclerite tapered posteriorly, connected to anterior sclerite by dorsal mound of membranous tissue.

Figures

FIGURE 1. Banyallarga (Banyallarga) argentinica. Adult (UMSP000022090): A—head, dorsal; B—thorax, dorsal.

FIGURE 2. Banyallarga (Banyallarga) argentinica. Wings (UMSP000022091): A—forewing; B—hindwing.

FIGURE 3. Banyallarga (Banyallarga) argentinica. Male (holotype): A—lateral view; B—terga VII–X, dorsal view; C—ventral view; D—phallus, lateral view; E—phallus, ventral view.

FIGURE 32. Banyallarga (Histricoverpa) sanchezi. Wings (holotype): A—forewing; B—hindwing.