Cymindis (Tarsostinus) rolandi Kabak et Schmidt, 2022

Kabak, I. I. & Schmidt, J., 2022, A new species of the genus Cymindis Latreille, 1805 (Coleoptera: Carabidae: Lebiini) from the Tarim River basin, China, Far Eastern Entomologist 446, pp. 1-6 : 3-6

Cymindis (Tarsostinus) rolandi Kabak et Schmidt , sp. n.

http://zoobank.org/NomenclaturalActs/ B010A35A-C3D8-44A6-8F60-AE2FE85A4E21

Figs 1–3

TYPE MATERIAL. Holotype – ♀, China: Xinjiang, Taklamakan, Ingbazar

(Tarimbrücke), 41°11' N, 84°14' E, h= 910 m, 29.VIII–3.IX 2004, leg. R. Schultz

( CJS). Paratype: 1 ♀, collected with holotype ( ZIN) .

DESCRIPTION. Female. Species medium-sized compared to other species of the subgenus, body length 9.6–9.7 mm, hind wings fully developed, habitus wide and subparallel-subconvex, appendages averaged ( Figs 1, 2 ). Colour of dorsal side:

head dark brown with reddish anterior part, labrum and mandibles; pronotum reddish brown with yellowish lateral sides; elytra brownish-yellow with dark sutural stripe

(on each elytron, 3–4 intervals in anterior half and 1–2 in apical third) and two symmetric longitudinal spots in posterior half of intervals 6–7 (either isolated or connected to sutural dark pattern). Legs monochromatic brownish yellow, antennae reddish. Ventral side of body yellowish-brown, head dark brown, lower surface densely punctate and pubescent.

Head rather large; eyes markedly protruding, glabrous, EyL/3AL = 1.14–1.20;

tempora long and flat, not delimited posteriorly by neck constriction. Antero-lateral margins of forehead significantly reflexed. Upper-side of head swollen at vertex,

frontal foveae hardly perceptible, supraorbital furrow very long, sharply and deeply impressed. Frons and vertex densely and roughly punctate. Pubescence of dorsal surface of head medium in length, moderately dense, slightly inclined forward. Two pairs of long supraorbital setae. Antennae comparatively short, EL/AL = 1.05–1.08.

Scape long, feebly constricted medially, sparsely pubescent, with long preapical seta.

Genae sparsely pubescent. Labial tooth shorter than lateral lobes, broadly rounded at apex, bordered along anterior margin. Submentum quadrisetose. Apical segment of labial palpi fusiform, with a few small hairs. Penultimate labial palpomere with four long setae on anterior margin. Apical maxillar palpomere fusiform, sparsely pubescent.

Pronotum unusually wide, PW/HW = 1.37–1.39, PW/PL = 1.50–1.58, markedly constricted toward base, PW/PB = 1.44–1.50, broadest in anterior third. Sides almost semicircular in anterior half, subrectilinear basally, shortly but sharply concave before laterobasal angles, the latter small, acute, slightly protruded laterally. Anterior margin deeply concave, without distinct border; anterior angles ample, markedly rounded and produced anteriorly. Basal margin of pronotum convex throughout,

bordered laterally. Lateral sides of pronotum extraordinarily explanate, especially in anterior half, and slightly reflexed. Disc subconvex medially, median line thin, significantly shortened anteriorly and posteriorly. Apical transverse impression distinct,

anterior surface markedly convex. Basal foveae shallow and vague, basal transverse impression deep, basal margin reflexed. Punctures rough and dense throughout pronotal surface, though smaller than those on head. Pubescence dense, suberrect, as long as on head. Two pairs of lateral setae present, one in anterior half of pronotum,

and one in laterobasal angles. Scutellum glabrous and smooth. Sides of prothorax spaciously pubescent. Metepisterna markedly longer than wide. Metacoxae, in addition to short pubescence, with two long setae.

view; 2 – the same, paratype; 3 – gonocoxites of paratype.

Elytra wide, subrectangular in shape, moderately convex, depressed on disc,

broadest in posterior half, EL/EW = 1.38–1.40, EL/PL = 3.00–3.01, EW/PW =

1.38–1.43. Lateral margins subrectilinear for most of their length, shoulders rounded, distinctly protruded anteriorly. Apices moderately oblique, slightly incised,

faintly ciliate; external apical angle of each elytron broadly rounded, sutural angle blunt. Marginal gutter moderate in width, narrowed near humeri, lateral margins slightly reflexed for most of their length. Basal border complete, markedly sinuate.

Elytral striae moderately deep, densely and faintly punctate. Both parascutellary striole and setiferous pores present. Intervals flat to subconvex, evenly and rather densely punctate, punctures arranged in 3–4 irregular rows; interval 3 with 3 small discal setiferous pores, of which the anterior one attached to stria 3. Pubescence of elytra dense, oblique, as long as on head and pronotum. Umbilicate series consists of

15–16 pores. One apical pore in stria 7 on the level either of stria 3 or of interval 3.

Microsculpture indistinct on head and pronotum, hardly perceptible to distinct on elytra, consisting of isodiametric meshes.

Visible abdominal sternites with a single pair of paramedian setae, anal sternite quadrisetose (females).

Meso- and metatibiae longer than corresponding tarsi. Dorsal surface of tarsi with long hairs; inner margin of claws mostly smooth, rarely with 1–4 very small denticles in basal quarter.

Apical gonocoxite rather narrow, moderately arched in distal half ( Fig. 3 ).

Male unknown.

COMPARATIVE DIAGNOSIS. The new species belongs to the subgenus Tarsostinus Motschulsky, 1864 due to the presence of denticles on inner claw margins,

two pairs of supraorbital setae and complete elytral basal border. Externally the new species ressembles C. ( Tarsostinus ) equestris Gebler, 1825 known from central and southern parts of Eastern Europe, Caspian Depression, Kazakh Upland, northern part of Middle Asia, Southern Siberia, Mongolia, eastern part of the Tibetan Plateau and adjacent plains (Emetz & Kryzhanovskij, 1973; Kabak, 2017). Both species share the fully developed hind wings, similar coloration, densely and roughly punctured body surface, deeply engraved supraorbital furrows, markedly explanate lateral sides of pronotum, very small to reduced denticles on inner margin of claws

(however, in the eastern populations of C. equestris claws are pectinate), and similar shape of the apical gonocoxite. The new species differs from C. equestris by its characteristic habitus: vertex swollen, pronotum markedly wider with much more broadly explanate lateral margins, less convex disc and reflexed anterior surface;

additionally, the eyes in the new species are glabrous (pubescent in C. equestris).

DISTRIBUTION. The new species is only known from the type locality: the flood plain of the Tarim River at the northern border of the Taklamakan Desert, to the south of Luntai Village, Xinjiang-Uygur Autonomous Region of China.

HABITAT. The new species was found in the end of summer in the flood plain of the Tarim river at an elevation of about 910 m a.s.l.

ETYMOLOGY. The new species is named after its collector, Dr. Roland Schultz,

specialist of ants (Görlitz, Germany).

We are very grateful to Dr. R. Schultz (Görlitz, Germany) for providing us with the carabid beetle material he collected during his expeditions to the Qinghai-Tibet

Plateau and the Tarim basin, and to Dr. G. Davidian (St-Petersburg, Russia) for his help in preparing photos.

Figures

Figs 1–3. Cymindis (Tarsostinus) rolandi sp. n., female. 1 – holotype, habitus, dorsal