Naticidae, Guilding, 1834

Kiel, Steffen, Fernando, Allan Gil S., Magtoto, Clarence Y. & Kase, Tomoki, 2022, Mollusks from Miocene hydrocarbon-seep deposits in the Ilocos-Central Luzon Basin, Luzon Island, Philippines, Acta Palaeontologica Polonica 67 (4), pp. 917-947 : 937-940

publication ID	https://doi.org/ 10.4202/app.00977.2022
persistent identifier	https://treatment.plazi.org/id/0470A855-EA6C-FF8E-FCAF-F905881CFE2E
treatment provided by	Felipe
scientific name	Naticidae
status

Treatment

Naticidae View in CoL gen. et sp. indet.

Fig. 20A View Fig .

Material.— One specimen from block B ( NRM Mo 192628: H = 21.3 mm, W = 21.2 mm). Upper Miocene of the “shale quarry” within the Northern Cement Corporation quarry in Pangasinan province, Luzon, Philippines .

Remarks.—Unfortunately, the umbilical area of this specimen is damaged, preventing a generic assignment.

Family Fasciolariidae Gray, 1853 View in CoL

Genus Fusinus Rafinesque, 1815 View in CoL

Type species: Murex colus Linnaeus, 1758 , by typification of replaced name; Recent, Indo-West Pacific Ocean.

Fusinus ? sp.

Fig. 20B, C View Fig .

Material.— Two large specimens from block I ( NRM Mo 192662, 192663). Upper Miocene of the “shale quarry” within the Northern Cement Corporation quarry in Pangasinan province, Luzon, Philippines .

Dimensions (in mm).— NRM Mo 192662: H = 73.2, W = 52.0; NRM Mo 192663: H = 90.8, W = 45.0.

Remarks.— Fusinus is widespread in the Indian and Pacific Oceans today ( Okutani and Tsuchiya 2000; Callomon and Snyder 2007, 2019; Couto et al. 2016), but its fossil record from this area is meager at best. Although the preservation of the two Fusinus ? sp. specimens is not ideal, they clearly lack the granulate sculpture typical for Granulifusus , a genus common in the Indo-West Pacific and known from several fossils, especially in Japan ( Shuto 1958; Hadorn and Fraussen 2005). With their evenly convex whorls, the two Fusinus ? sp. specimens are clearly different from those of several deep-water fusinins inhabiting deeper water around the Philippines today, such as species of Latirus Montfort, 1810 , and Fusolatirus Kuroda and Habe in Kuroda et al., 1971 ( Garcia 2001; Snyder and Bouchet 2006).

Vermeij and Snyder (2018) reviewed large fasciolariid genera (Fusininae) and introduced several genera for tall species with strongly convex whorls and distinct axial ribs resembling Fusinus ? sp. reported here. Among these, Apertifusus includes three or four tropical Atlantic species, and Lyonsifusus consists of a few species from the southern Caribbean, including Miocene fossils. Lastly, Ariefusus was introduced for an extant West African species and for the Miocene Fusus prevosti Partsch in Hörnes, 1856, whose whorls are more constricted near the suture ( Hörnes 1856) and consequently have somewhat pointed ribs on the center of the whorls, unlike the Philippine specimens.

Family Buccinidae Rafinesque, 1815 View in CoL

Genus Enigmaticolus Fraussen, 2008 View in CoL

Type species: Enigmaticolus monnieri Fraussen, 2008 , by original designation; Recent , Madagascar .

Remarks.—Six species and two unnamed records of this genus are known from the Indian and western Pacific Oceans Fraussen 2008; Fraussen and Stahlschmidt 2016; Zhang et al. 2020b; Chen et al. 2021). Here we transfer a species formerly reported as Siphonalia semisulcata Martin, 1933 , and Siphonalia (Pseudoneptunea) semisulcata , from the Upper Miocene of Buton, Indonesia, to Enigmaticolus , report it also from the Upper Miocene of the Philippines, and regard it as the first fossil record of this genus.

Enigmaticolus semisulcata ( Martin, 1933) comb. nov. Fig. 20D–F View Fig .

1933 Siphonalia semisulcata sp. nov.; Martin 1933: 25, pl. 3: 24, 25. 1942 Siphonalia (Pseudoneptunea) semisulcata Martin ; Beets 1942:

281, pl. 28: 38.

Description.—High-spired fusiform shell, whorls gently convex with fine constriction below suture; sculpture of flat spirals crossed by numerous fine axial (slightly opisthocyrt) growth increments, giving the spirals a beaded appearance; sculpture most prominent on early whorls and base; inner lip with thin callus.

Material.— Two specimens from block B ( NRM Mo 192586, 192587) , three specimens from block I ( NRM Mo 192631– 192633). Upper Miocene of the “shale quarry” within the Northern Cement Corporation quarry in Pangasinan province, Luzon, Philippines .

Dimensions (in mm).—NRM Mo 192586: H = 30.0, W = 21.5; NRM Mo 192586: H = 17.7, W = 11.8; NRM Mo 192632: H = 36.8, W = 24.0; NRM Mo 192632: H = 34.0, H = 21.0.

Remarks.—The specimens illustrated by Martin (1933) from the Upper Miocene of Buton, Indonesia, resemble the Philippine specimens reported here in general shape and in having flat, beaded spiral ornament that becomes more prominent on the base. They differ by having more distinctive axial ribbing in the upper part of the whorls, especially on the early whorls ( Martin 1933), whereas the Philippine specimens show more distinctive beaded spiral ornament. Even more similar to the Philippine specimens is another specimen from Buton, referred to as ” Siphonalia (Pseudoneptunea) semisulcata Martin ” by Beets (1942), due to its lack of the strong axial ribs on the upper part of the whorls.

Species of Siphonalia and Pseudoneptunea have more convex whorls with much stronger axial ornament than “ Siphonalia semisulcata Martin, 1933 ”; hence the species is unlikely to belong to either of these genera ( Makiyama 1941; Cernohorsky 1975; Robba et al. 2004; Zhang and Zhang 2018). Rather, the habitus of the shell, the flat spiral ornament on the upper part of the whorls and the base, the indistinct axial ribs, and fine constriction below the suture, indicate affinities to Enigmaticolus , to which this species is herein re-assigned. Among the extant species of Enigmaticolus , the most similar to E. semisulcata is the type species with its slender shell.

Stratigraphic and geographic range. —Upper Miocene of Luzon, Philippines and Buton, Indonesia.

Enigmaticolus sp.

Fig. 20G View Fig .

Material.— One specimen from block I ( NRM Mo 192640: H = 23.8 mm, W = 12.0 mm). Upper Miocene of the “shale quarry” within the Northern Cement Corporation quarry in Pangasinan province, Luzon, Philippines .

Remarks.—This specimen differs from Enigmaticolus semisulcata by being smaller and by having more distinct axial ribs on the whorl’s shoulder. In the latter respect it resembles Enigmaticolus nipponensis ( Okutani and Fujiwara, 2000) from methane seeps in the South China Sea ( Chen et al. 2021).