Haplodrassus bohemicus Miller & Buchar, 1977

Kovblyuk, Mykola M., Kastrygina, Zoya A. & Omelko, Mikhail M., 2012, A review of the spider genus Haplodrassus Chamberlin, 1922 in Crimea (Ukraine) and adjacent areas (Araneae, Gnaphosidae), ZooKeys 205, pp. 59-89 : 61-63

Haplodrassus bohemicus Miller & Buchar, 1977 Figs 1 –610– 12

Haplodrassus bohemicus Miller and Buchar 1977: 163, pl. II, f. 1-6 (♂♀).

Haplodrassus bohemicus : Stefanovska et al. 2008: 37, f. 10-15 (♂♀).

Material.

UKRAINE, CRIMEA: Saky Distr.: 12 ♂♂, 2 ♀♀ (TNU), near Pribrezhnaya railway station, 30.04.-24.06.2000, M.M. Kovblyuk.

Additional material.

UKRAINE. Nikolaev Area: 6 ♂♂ (TNU), Pervomaysky Distr., Kuripchane Vil., 5.05.-8.06.2006, N.Yu. Polchaninova. Kherson Area: 2 ♂♂, 2 ♀♀ (TNU), Henichesk Distr., Arabatskaya strelka, 4 km S Henichesk Town, 23.05.-10.06.2011, N.A. Stasyuk. Donetsk Area: 4 ♂♂ (TNU), Pershotravnevy Distr., Belosaraiskaya Kosa, 11-22.06.2001, E.V. Prokopenko; 2 ♂♂ (TNU), Novoazovsk Distr., Khomutovo Vil., "Khomutovskaya Steppe", N47°16', E38°10', 15-20.06.2004, N.Yu. Polchaninova; 1 ♂ (TNU), Slavyansky Distr., Svyatogorsk Town, N49°02', E37°39', 8-30.06.2004, N.Yu. Polchaninova. RUSSIA. Rostov Area: 7 ♂♂, 1 ♀ (TNU), Ust’ -Donetsk Distr., near Razdorskaya Vil., "Pukhlyakovskye sklony", 10.05.-28.06.2004, A.V. Ponomarev.

Comparative material.

Haplodrassus pugnans (Simon, 1880): RUSSIA, Magadan Area: 1 ♂, 3 ♀♀ (ISEA, БИ- 930), Ten’kynskyi Distr., env. Sibit-Tyellakh, Aborigen Field Station, Betula, 12-22.06.1983, A.V. Avershin.

Diagnosis.

Haplodrassus bohemicus is most similar to Haplodrassus pugnans (Simon, 1880), Haplodrassus signifer (C.L. Koch, 1839) and Haplodrassus pseudosignifer Marusiket al. 1996, but differs: 1) by the shape of RTA having a “step” -like dorsal margin (RTA dorsal margin without “step” in Haplodrassus pugnans , Haplodrassus signifer and Haplodrassus pseudosignifer ); 2) by the apically directed embolus lacking a tooth (retrolaterally directed embolus with a tooth in Haplodrassus pugnans , Haplodrassus signifer and Haplodrassus pseudosignifer ); 3) by the outlines of epigynal sclerites and relative proportions of the epigynal fovea (cf. Figs 10, 12 and 13, 75, 78).

Description.

Male measurements (n = 5). Total length 5.8-7.5 (6.7); carapace 2.7-3.2 (3.0) long, 2.0-2.6 (2.3) wide. Diameters of eyes and distances between them: AM 0.10-0.15 (0.12), AL 0.10-0.16 (0.13), PM 0.15-0.22 (0.19), PL 0.10-0.14 (0.12), AM-AM 0.09-0.12 (0.10), AM-AL 0.03-0.04 (0.04), PM-PM 0.03-0.04 (0.03), PM-PL 0.12-0.16 (0.14), AM-PM 0.12-0.18 (0.15), AL-PL 0.10-0.16 (0.14). Distances between anterior eyes and margin of clypeus: AM-clypeus 0.18-0.22 (0.20), AL-clypeus 0.10-0.18 (0.15).

Length of leg segments (male):

Length of palp segments: femur 1.0-1.2 (1.1), patella 0.4-0.5 (0.5), tibia 0.3-0.5 (0.4), tarsus 1.0-1.2 (1.0). Cheliceral teeth: anterior - 2 (little; proximal tooth connected with keel of cheliceral groove), posterior - 2. Abdomen 2.9-4.0 (3.6) long, 1.8-2.2 (2.0) wide. Scutum is absent. Basal segment of anterior (inferior) spinnerets 0.5-0.6 (0.6) long. Coloration light brown, as in most Haplodrassus species.

Palp as in Figs 1-6. RTA with a “step” (St) on the promargin, terminal apophysis almost straight, without distinct ridge, embolus slightly twisted and lacks a tooth.

Female measurements (n = 2). Total length 7.8-8.0; carapace 2.7-3.5 long, 2.0-2.7 wide. Abdomen 4.3-5.1 long, 2.7-3.1 wide. Coloration light brown, as in most Haplodrassus species.

Epigyne as in Figs 10-12. Lateral pockets long, sub-parallel, spermathecae globular. Fovea of epigyne without narrow longitudinal groove.

Distribution.

Czech Republic, Macedonia, Greece, Ukraine (Nikolaev, Kherson, Donetsk Areas, Crimea), Russia (Rostov and Stavropol Areas, Kalmykya, Dagestan) ( Miller and Buchar 1977; Ponomarev and Tsvetkov 2006; Stefanovska et al. 2008; Ponomarev et al. 2011; present data).

Comments.

Haplodrassus bohemicus is a new species record for the Crimean fauna.

Habitats.

Meadows, steppes and sand dunes.

Phenology.

♂♀ - V-VI. In Crimea the peak activity of adults occurs in May.