Ledomyia olgae Fedotova et Perkovsky, 2014

Fedotova, Z. A. & Perkovsky, E. E., 2014, New Gall Midges (Diptera, Cecidomyiidae, Brachineurini, Ledomyiini) From The Late Eocene Rovno Amber, Vestnik Zoologii 48 (1), pp. 35-50 : 48

Ledomyia olgae Fedotova et Perkovsky , sp. n.

M a t e r i a l. Holotype. SIZK N K–3697 , well preserved inclusion of male, Klesov , Rovno amber , Late Eocene ( fig. 6, 1–7 ; 8 , 3 , 4 ). Syninclusion: Diptera .

D e s c r i p t i o n. M a l e ( fig. 6, 1–7 ; 8 , 3 , 4 ). Body (without genitalia) is 1.3 times longer than antennae. Wings are 1.3 times as long as antennae. Antennae are 2+12-segmented. Head with large eyes that laterally cover only dorsal part of head. Scape is enlarged distally, almost as long as wide round pedicel. First flagellomere slightly longer than 2nd, its neck is shorter than the neck of the 2nd one; 5th flagellomere is 1.9 times as long as wide, its neck is 3.0 times shorter than basal enlargement. Tenth and 11th flagellomeres have very short neck; 10th flagellomere is 1.5 times as long as wide; 11th flagellomere is slightly narrowed distally and shorter than 10th. 12th flagellomere is oviform. Palpi are very narrow, 2–3-segmented, slightly swollen laterally. Fore tarsus with 2nd tarsomere 5 times as long as 1st. Wing is 2.2 times as long as wide. Vein R 1+2 joins C before its midlength; R 4+5 is strongly curved near apex, joins the wing margin closer to the apex than Cu 2. All tibiae subequal to or longer than respective femora. Fore tibia 1.2 times as long as femur. Hind leg is longer than fore and middle ones. Gonocoxites are slightly rounded laterally, 2.6 times as long as wide. Gonostylus is slightly curved, almost parallelsided, with wide apical claw, 1.7 times shorter than gonocoxites. Cerci are wider than gonocoxites. Aedeagus almost triangular.

M e a s u r e m e n t s, mm: body length, 0.76 (without genitalia); 0.83 (with genitalia); wing length, 0.77; wing width, 0.35; antennal length, 0.59; head length, 0.20 (without mouth parts); 0.25 (with mouth parts); width of lateral side of head, 0.11; palpi length, 0.12; thorax length, 0.28; abdomen length, 0.41; length of fore leg, 1.38; coxa and trochanter, 0.09; tarsus, 0.61; femur 0.32; tibia 0.36; 1st segment, 0.06, 2nd, 0.30; 3rd, 0.06; 4th, 0.09; 5th, 0.07; length of middle: leg, 1.31; femur 0.33; tibia 0.37; tarsus, 0.61; coxa and trochanter, 0.09; 1st segment, 0.06, 2nd, 0.28; 3rd, 0.12; 4th, 0.09; 5th, 0.06; length of hind: leg, 1.46; coxa and trochanter, 0.09; tarsus, 0.66; femur, 0.34; tibia, 0.37; 1st segment, 0.06, 2nd, 0.32; 3rd, 0.11; 4th, 0.10; 5th, 0.07.

C o m p a r i s o n. By the shape of the body, wing venation, long legs and form of flagellomeres the new species is closely related to Ledomyia dextra Fedotova et Perkovsky, 2008 from Dubrovitsa, differing from it by the 2+12-segmented antennae (vs. 2+11 of L. dextra ); wing ratio (2.2 vs. 2.6); very long 2nd tarsomere; wider and shorter middle flagellomeres with wide and short necks; all tibiae longer than respective femora (hind tibia of L. dextra shorter than femur); oviform apical flagellomere (vs. oval of L. dextra ); larger body size.

E t y m o l o g y. The species is named after Olga — daughter of E. E. Perkovsky.

All in all, in Rovno amber Brachineurini and Ledomyiini account for 11.9 % of all species of the superfamily Cecidomyioidea (27 vs. 227), in Baltic amber — for 9.0 % (6 vs. 67) ( Fedotova, Perkovsky, 2009).

Figures

Fig. 6. Ledomyia olgae, male, holotype (SIZK, N K–3697): 1 — general appearance; 2, 3 — palpi (variability of shape); 4 — flagellomeres 9–11; 5 — flagellomere 5; 6, 7 — genitalia (6 — ventral view,7 — dorsal view).Scale bars 0.1 mm.

Fig. 1. Rovnobrachineura kiryeyevi, male, holotype (SIZK, N K–3701): 1 — general appearance; 2 — scape, pedicel, flagellomeres 1–2; 3 — flagellomeres 8–10; 4 — flagellomeres 6–7; 5 — mouth parts and palpi; 6 — genitalia dorsally. Scale bars 0.1 mm.

Fig. 2. Brachineura polessica, female, holotype (SIZK, N K–1236): 1 — general appearance; 2 — scape, pedicel, flagellomeres 1–2; 3 — flagellomeres 9–10; 4 — flagellomeres 6–7; 5 — flagellomeres 3–5; 6 — palpus; 7 — middle tarsomeres 3–5; 8 — middle tarsomeres 1–2; 9 — middle tarsomere 1; 10 — middle tarsomere 5; 11 — ovipositor laterally. Scale bars 0.1 mm.

Fig. 3. Popovineura nacta, male, holotype (SIZK, N K–666): 1 — general appearance; 2 — scape, pedicel, flagellomeres 1–2; 3 — flagellomeres 5–6; 4 — flagellomeres 8–11; 5 — palpus; 6 — fore tarsus; 7 — fore tarsomere 5; 8 — genitalia. Scale bars 0.1 mm.

Fig. 4. Nascensluna mellea, female, holotype (SIZK, N K–1601): 1 — general appearance; 2 — flagellomeres 8–10; 3 — flagellomere 5; 4 — ovipositor. Scale bars 0.1 mm.

Fig. 5. Rhizomyia parkalovi, female, holotype (SIZK, N K–7385): 1 — general appearance; 2 — head; 3 — palpus; 4 — flagellomeres 9–10; 5, 6 — tarsomere 1 (variability of shape); 7 — flagellomere 5; 8 — apex of abdomen. Scale bars 0.1 mm.

Fig. 7. Rovnobrachineura kiryeyevi, male, holotype SIZK, N K–3701 (1–3); Brachineura polessica, female, holotype (SIZK, N K–1236) (4, 5); Rhizomyia parkalovi, female, holotype (SIZK, N K–7385) (6, 7): 1, 4, 6 — head and antenna (1 — ×129; 4 — ×62; 6 — ×117); 2 — genitalia (×283); 3, 5, 7 — general appearance (3 — ×46; 5 — ×29; 7 — ×39).

Fig. 8. Popovineura nacta, male, holotype (SIZK, N K–666) (1, 2); Ledomyia olgae, male, holotype (SIZK, N K–3697) (3, 4); Nascensluna mellea, female, holotype (SIZK, N K–1601) (5–7): 1–3, 7 — general appearance (1 — ×45; 2 — ×38; 3 — ×48; 7 — ×64); 4 — antennae (4 — ×129); 5 — hind leg (×82); 6 — head and antenna (×131).