Retoka hirsuta, Kondorosy, 2008

Kondorosy, Előd, 2008, A revision of the Entisberus group (Hemiptera: Heteroptera: Rhyparochromidae), Acta Entomologica Musei Nationalis Pragae 48 (2), pp. 591-610 : 598-599

Retoka hirsuta sp. nov.

( Fig. 5 , 16 )

Type locality. Philippines, Luzon, Lagunas, Mt. Makiling, above Mad Springs.

Type material. HOLOTYPE: J, ‘ Philippines, Luzon : Lagunas , / Mt. Makiling, above Mad Springs, / 400-700m, degrad. rainforest, / 19.-22. XI. 1995, J. Kodada leg.’ ( NHMW) . PARATYPES: J, ‘ Malaysia, Sabah, / Crocker Range , 17. 6. 1996, / Mawar Waterfall env. (9c) // vegetation debris and / forest litter around / fallen trees’ ( NHMW) ; J, ‘MALAYSIA-SE SABAH nr. / Darum Valley Field C. / E1, c. 150m, Mal. trap 7 / 15-21. III. 1987, RMNH / C. v. Achterberg’ ( RMNH) .

Description. Macropterous male. Structure. Body oval, shining, with erect and sparse pilosity. Hairs on dorsum about twice as long as diameter of antennal segment I. Hairs on antennae partly erect, as long as hairs of dorsum of body, partly semi-decumbent, as long as diameter of antennal segments.

Head triangulate with eyes slightly pedunculate, slightly removed from anterior margin of pronotum.Antennal segments spindle-shaped, segment I surpassing apex of clypeus by more than half of its length. Labium reaching mid coxae, segment I reaching about half of length of head, thicker than following segments. Bucculae relatively high, of even height, reaching almost base of head, posteriorly pointed.

Pronotum trapezoidal, anterior part evenly convex. Anterior margin with distinctly punctate collar. Anterior and posterior parts of pronotum clearly separated by a distinct transverse impression. Anterior margin straight, posterior margin slightly sinuate anterior of scutellum and strongly sinuate anterior of both clavi. Lateral margins of pronotum concave at transverse impression, narrowly and evenly laminate, abruptly terminating at base. Calli impunctate except a few punctures, pronotum strongly punctate between calli. Posterior half of pronotum with broad and shallow median furrow. Posterolateral angles each with impunctate elevation. Scutellum triangular, with elevated impunctate Y-shaped carina; elevation almost as high as posterior part of pronotum; anterior arms well developed, reaching to anterolateral angles of scutellum. Clavus with three indistinctly separated rows of strong punctures. Corium with two rows of punctures along claval suture, endocorium with triangular impunctate area adjacent to the rows, meso- and exocorium evenly punctate. Basal half of costal margin of corium and lateral margin of pronotum with minute teeth.Apical margin of corium sinuate. Mesosternum with smooth depression posterior to fore coxae, delimited with strong tooth-like elevation laterally. Metasternum smooth, with transversal row of punctures at middle. Metasternal scent gland area small, not occupying more than one third of metasternum.

Abdomen. Sutures between abdominal sternites densely striolate longitudinally. Femora unarmed, tibiae straight.

Male genitalia. Paramere thin, with moderately acute lobes, distal part long ( Fig. 16 ).

Colouration. Body dark brown, head (except clypeus) blackish, antennae light brown, clavus, basal part of corium to level of end of scutellum, rostrum, and legs stramineous, membrane translucent.

Measurements (all in mm; holotype J first, paratypes (2JJ) in parentheses). Total body length 2.32 (2.18-2.22). Head: length 0.45 (0.42), width 0.65 (0.60), interocular space 0.38 (0.32-0.35). Pronotum: length 0.52 (0.50-0.52), width at base 1.08 (1.05-1.08). Scutellum length 0.48 (0.48), length of claval commissure 0.15 (0.18-0.20). Lengths of antennal segments I 0.38 (0.32-0.38), II 0.38 (0.35-0.40), III 0.28 (0.28-0.30), IV 0.35 (0.35). Lengths of labial segments of holotype I 0.28, II 0.35, III 0.18, IV 0.20.

Differential diagnosis. Within Retoka , this species is most dissimilar to the type species, R. minuta . Its unique characters include slightly pedunculate eyes, body covered with long hairs, well visible and very long bucculae that, almost reach the base of the head, well-developed anterior arms of the Y-shaped scutellar carina, and a depressed mesosternum.

Etymology. The species epithet is the Latin adjective hirsutus (= hairy), referring to the hairy body of this species.

Distribution. Philippines (Luzon), Malaysia (Borneo: Sabah).

Figures

Figs. 5-8. Habitus of Retoka species, body in dorsal view. 5 – R. hirsuta sp. nov.; 6 – R. gibba (Zheng, 1981); 7 – R. esakii (Slater & Hidaka, 1958); 8 – R. callosa (Scudder, 1968).

Figs. 14-21. Parameres. 14 – Entisberus archetypus Distant, 1903 (specimen from Vietnam: Yen Bai Province); 15 – E. bergrothi sp. nov.; 16 – Retoka hirsuta sp. nov,; 17 – R. gibba (Zheng, 1981); 18 – R. esakii (Slater & Hidaka, 1958); 19 – R. curvipes sp. nov.; 20 – R. stysi sp. nov.; 21 – R. paraminuta sp. nov.