Retoka minuta China , 1935

Kondorosy, Előd, 2008, A revision of the Entisberus group (Hemiptera: Heteroptera: Rhyparochromidae), Acta Entomologica Musei Nationalis Pragae 48 (2), pp. 591-610 : 604-608

Retoka minuta China, 1935

( Fig. 12 )

Retoka minuta China, 1935: 302 . Type locality: Sumatra, Tobagebiet Hochfläche von Huta Gindjang. Retoka minuta: SLATER (1964) : 950 (catalogue).

Type material examined. HOLOTYPE: ♀, Sumatra, Tobagebiet Hochfläche von Huta Gindjang ( BMNH). The holotype is in bad condition: it is partially broken with the abdomen, left hemelytron, and legs partly missing. However, it is possible to identify additional specimens by direct comparison . Additional material examined. TAIWAN: ‘Formosa, Pilam, ii.1908., Sauter leg.’ (♀, HNHM).

Distribution. Indonesia (Sumatra) ( CHINA 1935), Taiwan. New species for the fauna of Taiwan.

Retoka paraminuta sp. nov.

( Fig. 13 , 21 )

Type locality. India, West Bengal, Darjeeling District, Goomti.

Type material. HOLOTYPE: J, ‘ INDIA, W Bengal / Darjeeling Distr. / Goomti, 1200 m / leg. Gy. Topál // netting in under- / growth / 27.v.1980 ’. ( HNHM).

Description. Structure. Body oval, shining; dorsum covered by very minute hairs, sparsely distributed long erect hairs present on pronotum and hemelytra; body except abdomen very densely and strongly punctate.

Head short, about 0.8 times as long as high, moderately declivent at base of clypeus in lateral view, very densely punctate (having the appearance of very narrow but high mesh-like carinae). Eyes large, bare, strongly protruding but not pedunculate, laterally far surpassing anterolateral angles of pronotum, projecting posteriad. Posterior margin of eyes approaching pronotum but not touching it. Ocelli well developed, adjacent with pronotum. Base of antenna close to eye (distance between anterior margin of eye and base of antenna a little longer than half of eye length), projecting inwards. Bucculae small, rounded, very narrow posteriorly, reaching anterior margin of eyes. Antennal segments moderately thickened, with decumbent and prominent hairs; segment I surpassing apex of head by one-third of its total length; segment II longer than segment I; remaining segments shorter than segment II. Labium short, reaching apices of anterior coxae, segment I reaching about middle of eyes, segment II subequal in length to segment I, remaining segments shorter than segments I and II.

Thorax. Pronotum convex, only slightly concave at transverse furrow, anterior part very densely and posterior lobe slightly more sparsely punctate. Pronotum with small elevated impunctate areas on posterior lobe, anterior lobe with some impunctate areas, humeral angles smooth. Pronotal collar indistinct, with two irregular rows of punctures. Transverse impression shallow but well visible, situated at middle of pronotum. Lateral margins narrowly explanate, not laminate at level of transverse impression, abruptly terminating posteriorly, reaching collar anteriorly. Basal margin of pronotum protruding posteriad in front of scutellum, laterally depressed, apical margin straight. Scutellum triangular, with well-developed Y-shaped elevation, its median part high, reaching level of pronotum, reaching neither basal nor apical angles, basal arms strong, directed posteriad towards basal angles, apical arm smooth; middle of elevation behind one-third of total length of scutellum. Hemelytra pseudobrachypterous, reaching end of abdomen. Clavus narrow, with three regular rows of punctures. Corium elevated, highest near end of clavus, with two rows of punctures along claval margin, smooth lateral of them to corial furrow, mesocorium with dense irregular punctures, costal margin very narrow, impunctate. Costal margin of corium convex (concavity hardly visible at end of scutellum). Apical corial margin S-shaped, inner part only slightly concave. Membrane with four well visible veins. Sternum evenly punctate. Meso- and metasternum with prominent median ridge. Metasternal scent gland area small, not occupying one third of metasternum. Legs slender, fore femur with 4-5 fine teeth, fore tibia always straight and simple.

Abdominal venter without median keel. Intersegmental sutures between sternites II-VI punctate.

Male genitalia. Paramere thick with widely rounded outer lobe ( Fig. 21 ).

Colouration. Body dark brown. Head fuscous, tibiae, antennal segment I, and abdomen reddish brown. Posterior pronotal lobe, antennal segments II-IV, scutellum, and hemelytra yellowish brown. Pronotal lateral carinae, apical arm of scutellar carina, two obscure spots on costal margin of hemelytra, tarsi, and labium stramineous. Membrane whitish.

Measurements (all in mm;J holotype). Total body length 1.60, total width (across humeri) 0.85. Head: length 0.30, width 0.50, interocular space 0.28. Pronotum: length 0.45, width at base 0.75. Scutellum length 0.32, length of claval commissure 0.20. Lengths of antennal segments I 0.20, II 0.20, III 0.18, IV 0.25. Lengths of labial segments I 0.18, II 0.18, III 0.10, IV 0.15.

Differential diagnosis. The smallest species of the genus, distinctly smaller than all other species except of R. minuta and R. stysi sp. nov., which are only slightly larger than R. paraminuta sp. nov. Retoka minuta differs from the new species mainly in the following characters: hemelytra semicircular; body without long hairs; clavus wide, apical part with a few additional punctures between the two inner rows of punctures; corium totally punctate, anterior pronotal lobe without impunctate elevations. Retoka stysi sp. nov. differs from R. paraminuta sp. nov. in the following characters (character states of R. paraminuta sp. nov. given in parentheses): anterior pronotal lobe and first antennal segment black (brown), second antennal segment brown (yellow); clavus with some additional punctures outside the rows (without additional punctures); scutellar elevation very high in the middle, surpassing level of pronotum, anterior part steeper than 45° in lateral view (basal arms only slightly lower than medial part, elevation not surpassing level of pronotum, anterior part never steeper than 45° in lateral view).

Etymology. The species epithet refers to the strong resemblance to R. minuta .

Distribution. India (West Bengal).

Figures

Figs. 9-12. Habitus of Retoka species (9, 11-12 – body in dorsal view; 10 – detail of right anterior leg of male holotype). 9-10 – R. curvipes sp. nov.; 11 – R. stysi sp. nov.; 12 – R. minuta China, 1935.

Fig. 13. Habitus of Retoka paraminuta sp. nov.

Figs. 14-21. Parameres. 14 – Entisberus archetypus Distant, 1903 (specimen from Vietnam: Yen Bai Province); 15 – E. bergrothi sp. nov.; 16 – Retoka hirsuta sp. nov,; 17 – R. gibba (Zheng, 1981); 18 – R. esakii (Slater & Hidaka, 1958); 19 – R. curvipes sp. nov.; 20 – R. stysi sp. nov.; 21 – R. paraminuta sp. nov.