Temnothorax rugosus ( Mackay, 2000 )

Prebus, Matthew M., 2021, Taxonomic revision of the Temnothorax salvini clade (Hymenoptera: Formicidae), with a key to the clades of New World Temnothorax, PeerJ (e 11514) 9, pp. 1-462 : 351-358

Temnothorax rugosus ( Mackay, 2000)

Distribution: Fig. 150B ; worker, gyne & variability: Fig. 152 .

Leptothorax rugosus Mackay, 2000: 399 , figs. 18, 160. Worker. Nuevo León, Mexico.

Temnothorax rugosus (Mackay) : Bolton, 2003: 272. First combination in Temnothorax .,

Type material examined: Paratype worker: MEXICO: Nuevo León: El Salto (Zaragosa), 1,415 m, 10 June 1988, W. Mackay #WM11012-10 (images of UTEPENT05451 examined on antweb.org) [ UTEP] .

Non-type material examined: MEXICO: Hidalgo: 11 km west of Jacala , 1,585 m, 14 January 1952, W.S. Creighton, in Quercus polymorpha , 3 workers, 1 dealate gyne (LACMENT323267) [ LACM] 4 workers (LACMENT323268) [ LACM] 4 workers (LACMENT323269) [ LACM] 4 workers (LACMENT323270) [ LACM] 4 workers (LACMENT323272) [ LACM] 3 workers (LACMENT323333) [ LACM] ; same data as previous, except: in oak twig, 3 workers (LACMENT323280) [ LACM] 2 workers (LACMENT323324) [ LACM] 3 workers, 1 dealate gyne (LACMENT323326) [ LACM] 3 workers, 1 dealate gyne (LACMENT323327) [ LACM] 3 workers, 1 dealate gyne (LACMENT323328) [ LACM] 3 workers, 1 dealate gyne (LACMENT323271) [ LACM] ; same data as previous, except: in oak limb, 4 workers, 1 dealate gyne (LACMENT323329) [ LACM] 4 workers, 1 dealate gyne (LACMENT323330) [ LACM] . Nuevo León: Monterrey, Chipinque Mesa, 1,280 m, 21 January 1952, W.S. Creighton, in Quercus clivicola , 3 workers (LACMENT323302) [ LACM] 4 workers (LACMENT323303) [ LACM] 4 workers (LACMENT323306) [ LACM] 4 workers (LACMENT323307) [ LACM] 4 workers (LACMENT323314) [ LACM] 4 workers (LACMENT323325) [ LACM] ; same data as previous, except: in Quercus canbyi , 4 workers (LACMENT323308) [ LACM] 4 workers (LACMENT323309) [ LACM] 4 workers (LACMENT323310) [ LACM] 4 workers (LACMENT323311) [ LACM]; 22 January 1952, in Quercus clivicola , 3 workers, 1 dealate gyne (LACMENT323286) [ LACM] 4 workers (LACMENT323287) [ LACM] 4 workers (LACMENT323288) [ LACM] 4 workers (LACMENT323289) [ LACM] 4 workers (LACMENT323290) [ LACM] 4 workers (LACMENT323291) [ LACM] ; same data as previous, except: 1,280 m, 18 February 1952, in Quercus canbyi , 4 workers (LACMENT323297) [ LACM] 4 workers (LACMENT323298) [ LACM] 4 workers (LACMENT323299) [ LACM] 4 workers (LACMENT323300) [ LACM] 4 workers (LACMENT323301) [ LACM] 4 workers (LACMENT323304) [ LACM] ; same data as previous, except: 975 m, 19 February 1952, in Quercus sp. , 3 workers, 1 alate gyne (LACMENT323337) [ LACM] 4 workers (LACMENT323334) [ LACM] 4 workers (LACMENT323335) [ LACM] 4 workers (LACMENT323336) [ LACM] 4 workers (LACMENT323338) [ LACM] ; same data as previous, except: in Quercus clivicola , 3 workers, 1 dealate gyne (LACMENT323273) [ LACM] 4 workers (LACMENT323274) [ LACM] 3 workers, 1 dealate gyne (LACMENT323276) [ LACM] 4 workers (LACMENT323279) [ LACM] ; same data as previous, except: in Quercus canbyi , 4 workers (LACMENT323275) [ LACM] 4 workers (LACMENT323277) [ LACM] 3 workers (LACMENT323278) [ LACM] ; same data as previous, except: 1,100 m, in Cercis reniformis , 4 workers (LACMENT323292) [ LACM] 4 workers (LACMENT323293) [ LACM] 4 workers (LACMENT323294) [ LACM] 4 workers (LACMENT323295) [ LACM] 4 workers (LACMENT323296) [ LACM] 4 workers (LACMENT323361) [ LACM] ; same data as previous, except: 1,220 m, in Quercus canbyi , 3 workers (LACMENT323281) [ LACM] 4 workers (LACMENT323282) [ LACM] 4 workers (LACMENT323283) [ LACM] 4 workers (LACMENT323284) [ LACM] 4 workers (LACMENT323285) [ LACM] 3 workers, 1 dealate gyne (LACMENT323322) [ LACM] ; same data as previous, except: 1,280 m, 21 April 1952, in Quercus clivicola , (LACMENT323305) [ LACM] 4 workers (LACMENT323318) [ LACM] 4 workers (LACMENT323319) [ LACM] 4 workers (LACMENT323320) [ LACM] ; same data as previous, except: in Quercus canbyi , 4 workers (LACMENT323312) [ LACM] 4 workers (LACMENT323313) [ LACM] 4 workers (LACMENT323315) [ LACM] 4 workers (LACMENT323316) [ LACM] 4 workers (LACMENT323317) [ LACM] 4 workers (LACMENT323321) [ LACM] 4 workers (LACMENT323323) [ LACM]; 10 km west of Iturbide, 1,890 m, 16 February 1952, W.S. Creighton, in Quercus clivicola , 4 workers (LACMENT323358) [ LACM] 4 workers (LACMENT323360) [ LACM] 4 workers (LACMENT323362) [ LACM] 4 workers (LACMENT323363) [ LACM] 4 workers (LACMENT323364) [ LACM] 4 workers (LACMENT323365) [ LACM] 4 workers (LACMENT323366) [ LACM] 3 workers, 1 dealate gyne (LACMENT323367) [ LACM]; 3 km west of Iturbide , 1,675 m, 16 February 1952, W.S. Creighton, in hollow stem of Rhus sp. , 4 workers (LACMENT323359) [ LACM] .

U.S.A.: Texas: Jeff Davis County: Davis Mountains State Park , 1,525 m, 11 May 1953, W.S. Creighton, in Quercus grisea , 3 workers (LACMENT323257) [ LACM] ; same data as previous, except: in deciduous oak, 2 workers (LACMENT323258) [ LACM] ; same data as previous, except: 16 May 1953, W.S. Creighton, in Quercus grisea , 3 workers (LACMENT323252) [ LACM] 2 workers (LACMENT323253) [ LACM] 4 workers (LACMENT323254) [ LACM] 3 workers (LACMENT323255) [ LACM] 3 workers (LACMENT323256) [ LACM] 3 workers, 1 dealate gyne (LACMENT323259) [ LACM] 4 workers (LACMENT323260) [ LACM] 4 workers (LACMENT323261) [ LACM] 4 workers (LACMENT323262) [ LACM] 4 workers (LACMENT323263) [ LACM] ; same data as previous, except: 20 May 1952, W.S. Creighton, in Quercus grisea , 4 workers (LACMENT323264) [ LACM] 4 workers (LACMENT323265) [ LACM] 4 workers (LACMENT323266) [ LACM] 3 workers (LACMENT323331) [ LACM] 2 workers (LACMENT323332) [ LACM] 3 workers (LACMENT323339) [ LACM] .

Geographic range: Mid-to-high elevations of southwestern Texas to Hidalgo State, Mexico ( Fig. 150B ).

Worker diagnosis: Temnothorax rugosus can be separated from all other species in the salvini clade by the following character combination: anterior clypeal margin emarginate; metanotal groove absent; propodeum not depressed; hind femora weakly to strongly incrassate; petiole cuneiform to subcuneiform; dorsum of head, mesosoma, waist segments and gaster with erect, short, blunt-tipped setae; legs without erect setae; integument testaceous yellow, with legs lighter yellow.

Similar species: Temnothorax acuminatus sp. nov., T. nitens , T. tricarinatus , species of the annexus and silvestrii groups. Temnothorax rugosus may be separated from T. nitens and T. tricarinatus by the moderately incrassate femora (not apparent in smaller workers), and the weakly sculptured first gastral tergite (which is smooth and shining in T. nitens and T. tricarinatus ). Many of the preceding characters, in combination, are shared with members of the annexus species group, which T. rugosus is closely related to. Temnothorax rugosus can be separated from the species of the annexus group by the cuneiform petiolar node, which is laterally compressed in dorsal view; all species in the annexus group have quadrate to subquadrate petiolar nodes. Finally, the areolate first gastral tergite is similar to members of the silvestrii group, but members of that group never have incrassate femora, and always have subquadrate petiolar nodes.

Worker measurements & indices (n = 27): SL = 0.465 –0.602 (0.527); FRS = 0.207 –0.279 (0.228); CW = 0.610 –0.800 (0.700); CWb = 0.563 –0.733 (0.642); PoOC = 0.265 –0.325 (0.288); CL = 0.701 –0.874 (0.773); EL = 0.157 –0.255 (0.198); EW = 0.116 –0.166 (0.142); MD = 0.173 –0.217 (0.197); WL = 0.824 –1.095 (0.948); SPST = 0.141 –0.219 (0.182); MPST = 0.203 –0.334 (0.281); PEL = 0.242 –0.362 (0.291); NOL = 0.147 –0.230 (0.178); NOH = 0.105 –0.180 (0.133); PEH = 0.226 –0.303 (0.260); PPL = 0.170 –0.289 (0.211); PPH = 0.210 –0.306 (0.244); PW = 0.368 –0.502 (0.428); SBPA = 0.135 –0.213 (0.160); SPTI = 0.148 –0.241 (0.187); PEW = 0.149 –0.217 (0.178); PNW = 0.066 –0.117 (0.090); PPW = 0.245 –0.393 (0.290); HFL = 0.484 –0.680 (0.578); HFWmax = 0.148 –0.209 (0.177); HFWmin = 0.049 –0.081 (0.059); CS = 0.914 –1.170 (1.029); ES = 0.219 –0.327 (0.269); SI = 77–89 (82); OI = 24–29 (26); CI = 80–87 (83); WLI = 139–157 (148); SBI = 22–29 (25); PSI = 17–22 (19); PWI = 140–181 (163); PLI = 113–176 (139); NI = 105–173 (135); PNWI = 38–67 (50); NLI = 49–75 (61); FI = 225–354 (301).

Worker description: In full-face view, head subquadrate, longer than broad (CI 80–87). Mandibles densely, finely striate but shining and armed with five teeth: the apical-most well developed and acute, followed by a less developed preapical tooth and three equally developed smaller teeth. Anterior clypeal margin emarginated medially. Antennal scapes short: when fully retracted, failing to reach the posterior margin of the head capsule by about two times the maximum width of the antennal scape (SI 77–89). Antennae 12-segmented; antennal club of composed of three segments, with the apical-most segment about one and a half times as long as the preceding two in combination. Frontal carinae short, extending past the antennal toruli by about two times the maximum width of the antennal scape. Compound eyes moderately protruding past the lateral margins of the head capsule. margin of head weakly convex, forming a continuous arc from the mandibular insertions to the posterior margin of the head. Posterior head margin concave medially, rounding evenly into the lateral margins.

In profile view, compound eyes ovular and moderately large (OI 24–29), with 13 ommatidia in longest row. Pronotal declivity distinct: dorsal margin of anterior face of pronotum marked by a weak carina and change in sculpture; neck and anterior face of pronotum forming a ~120 ° angle. Mesosoma very weakly sinuate: promesonotum forming an even convexity from where it joins the pronotal declivity; propodeum weakly depressed below the level of the promesonotum and descending sharply to the base of the propodeal spines on the posterior quarter. Promesonotal suture extending from the posterior margin of the procoxal insertion to the mesothoracic spiracle, which is well developed. Metanotal groove visible as a disruption of the sculpture laterally from where it arises between the mid- and hind coxae to where it ends in the poorly developed metathoracic spiracle, which is nearly indistinguishable against the ground sculpture. Propodeal spiracle moderately well developed, directed posterolaterally, and separated from the propodeal declivity by about four spiracle diameters. Propodeal spines weakly developed and short (PSI 17–22), about a third as long as the propodeal declivity, flared at the base, stout, triangular, and acute. Propodeal declivity straight, forming a rounded ~120 ° angle with the base of the propodeal spines. Propodeal lobes rounded and weakly developed. Metapleural gland bulla small, extending from the metacoxal insertion a third of the way to the propodeal spiracle. Petiole moderately long (PLI 113–176), with tubercles anterodorsally. Subpetiolar process in the form of a blunt, triangular tooth which grades evenly into the ventral petiole margin posteriorly. Petiolar peduncle short: comprising about a quarter of the total petiole length. Petiolar node truncate but dorsum is extremely angled, so that in many specimens it appears cuneiform: anterior face forming a sharp ~90 ° angle with the dorsal face, which is flat; dorsal face meeting the posterior face at a sharp ~120 ° angle; very short posterior face forms a ~110 ° angle with the caudal cylinder. Postpetiole evenly convex anterodorsally; weakly lobed ventrally.

In dorsal view, humeri developed: angulate anterolaterally and slightly wider than the rest of the mesosoma; mesothoracic spiracles protruding past the lateral margins of the mesosoma, visible as slight angles where the pronotum meets the mesonotum. Promesonotal suture represented by a weak sulcus and disruption in the ground sculpture. Metanotal groove absent: mesonotum and propodeum completely fused and lateral margins converging evenly to the bases of the propodeal spines. Propodeal spines broadly approximated basally and diverging apically, their apices separated from each other by about twice their length, the negative space between them “U” shaped. Petiolar peduncle with spiracles weakly protruding past the lateral margins. Petiolar node, when viewed at posterodorsal angle, cuneiform: narrowed apically and flat dorsally; node narrower than the peduncle and the caudal cylinder. Postpetiole narrow (PWI ~ 163) and campaniform. Anterior margin of the postpetiole convex and rounds evenly into the lateral margins; lateral margins parallel to each other; posterior corners rounded; posterior margin flat. Metafemur moderately to strongly incrassate (FI 225–354).

Sculpture: median clypeal carina present, extending posteriorly to the frontal triangle, and flanked on either side by three weaker carinae. Lateral clypeal lobes with additional, weaker carinae; ground sculpture shining through weak areolae. Antennal scapes areolate. Cephalic dorsum areolate, with costa over the ground sculpture, which become rugose on the posterior sixth of the head; concentric rugulae surrounding the antennal insertions. Lateral surfaces of head areolate, with coarse rugae over the ground sculpture. Ventral surface of head smooth and shining medially, but otherwise weakly areolate with weak costulae. Pronotal neck areolate. Lateral surfaces of mesosoma weakly areolate, with coarse, costae over the ground sculpture; region between the propodeal spiracle and propodeal spines areolate, without rugae. Propodeal declivity areolate, with fine strigulae. Dorsal surface of mesosoma weakly areolate, with coarse costae that are rugose on the pronotum and the propodeum. Femora finely, weakly areolate. Petiole uniformly areolate; a weak carina present laterally, extending longitudinally from the petiolar spiracle to the caudal cylinder; rugae on lateral faces of the node. Postpetiole uniformly areolate, with rugose sculpture on the dorsal and lateral surfaces. First gastral tergite shining through weak areolate sculpture; without spectral iridescence. First gastral sternite smooth and shining, without spectral iridescence.

Setae: antennal scapes and funiculi with short, subdecumbent pilosity. Dorsum of the head, pronotum, waist segments, and gaster with moderately abundant, erect, blunt-tipped setae, the longest of which are about half the width of the compound eye. The head bears ~50, mesosoma ~26, petiole 8, postpetiole ~32, and first gastral tergite ~72 setae. Short, sparse pubescence present over the entire body, but difficult to detect against the densely sculptured integument.

Color: predominantly testaceous yellow, with legs lighter yellow.

Gyne measurements & indices (n = 5): SL = 0.518 –0.600 (0.556); FRS = 0.264 –0.309 (0.291); CW = 0.783 –0.831 (0.802); CWb = 0.726 –0.782 (0.746); PoOC = 0.304 –0.319 (0.312); CL = 0.824 –0.880 (0.851); EL = 0.233 –0.277 (0.253); EW = 0.171 –0.207 (0.191); MD = 0.169 –0.196 (0.183); WL = 1.324 –1.477 (1.401); SPST = 0.202 –0.233 (0.223); MPST = 0.322 –0.383 (0.354); PEL = 0.312 –0.382 (0.355); NOL = 0.204 –0.228 (0.213); NOH = 0.145 –0.168 (0.155); PEH = 0.300 –0.351 (0.319); PPL = 0.215 –0.238 (0.224); PPH = 0.314 –0.376 (0.345); PW = 0.783 –0.838 (0.802); SBPA = 0.300 –0.391 (0.338); SPTI = 0.268 –0.340 (0.310); PEW = 0.222 –0.269 (0.237); PNW = 0.120 –0.185 (0.146); PPW = 0.378 –0.465 (0.413); HFL = 0.644 –0.718 (0.667); HFWmax = 0.166 –0.189 (0.178); HFWmin = 0.053 –0.069 (0.062); CS = 1.138 –1.216 (1.171); ES = 0.319 –0.381 (0.349); SI = 71–77 (75); OI = 28–33 (30); CI = 85–90 (88); WLI = 182–200 (188); SBI = 41–52 (45); PSI = 15–17 (16); PWI = 165–186 (174); PLI = 145–176 (158); NI = 121–146 (138); PNWI = 52–79 (62); NLI = 53–67 (60); FI = 242–313 (289).

Gyne description: In full-face view, head subquadrate, longer than broad (CI 85–90). Mandibles densely striate but shining and armed with five teeth: the apical-most well developed, followed by a less developed preapical tooth and three equally developed smaller teeth. Anterior clypeal margin emarginated medially. Antennal scapes short: when fully retracted, failing to reach the posterior margin of the head capsule by about two times the maximum width of the scape (SI 71–77). Antennae 12-segmented; antennal club composed of three segments, with the apical-most segment about one and a half times as long as the preceding two in combination. Frontal carinae moderately long, extending past the antennal toruli by about two and a half times the maximum width of the antennal scape. Compound eyes strongly protruding past the lateral margins of the head capsule. Lateral margins of head convex behind the compound eyes, but parallel to each other between the compound eyes and the mandibular insertions. Posterior head margin weakly concave medially, rounding evenly into the lateral margins.

In profile view, compound eyes ovular and large (OI 28–33), with 17 ommatidia in longest row. Mesoscutum rounded evenly anteriorly, covering the dorsal surface of the pronotum, and flat dorsally. Mesoscutellum on the same plane as the mesoscutum and flat dorsally. Posterior margin of metanotum extending slightly past the posterior margin of the mesoscutum. Propodeal spiracle well developed, directed posterolaterally, and separated from the propodeal declivity by about four spiracle diameters. Propodeal spines stout, weakly developed, and short (PSI 15–17), about a third as long as the propodeal declivity, flared at the base, triangular, and blunt. Propodeal declivity straight and flat, forming a rounded ~120 ° angle with the base of the propodeal spines. Propodeal lobes rounded. Metapleural gland bulla small, extending from the metacoxal insertion halfway to the propodeal spiracle. Petiole moderately long (PLI 145–176), with tubercles anterodorsally. Subpetiolar process in the form of a blunt, triangular tooth, which grades evenly into the ventral margin of the petiole posteriorly. Petiolar peduncle short: comprising about a quarter of the total petiole length. Petiolar node truncate but dorsum is extremely angled, so that in many specimens it appears cuneiform: anterior face forming a sharp ~90 ° angle with the dorsal face, which is flat; dorsal face meeting the posterior face at a sharp ~120 ° angle; very short posterior face forms a ~110 ° angle with the caudal cylinder. Postpetiole evenly convex anterodorsally, flat posterodorsally; weakly lobed ventrally.

In dorsal view, mesoscutum covering pronotum anteriorly, but humeri visible laterally as rounded sclerites. Propodeal spines weakly diverging apically, their apices separated from each other by about three times their length. Petiolar peduncle with spiracles weakly protruding past the lateral margins. Petiolar node, when viewed posterodorsally, tapering apically so the apex is about half the width of the base; dorsal margin emarginate medially. Postpetiole narrow (PWI 165–186), anteroposteriorly compressed, and subquadrate. Anterior margin of postpetiole weakly convex but emarginated medially; anterior corners marked by rounded ~100 ° angles as it transitions to the lateral margins; lateral margins parallel to each other; posterior corners rounded; posterior margin weakly concave. Metafemur weakly to strongly incrassate (FI 242–313).

Sculpture: median clypeal carina present, extending from the anterior margin nearly to frontal triangle, and flanked on either side by three equally strong carinae. Lateral clypeal lobes with additional weaker carinae; ground sculpture weakly areolate. Antennal scapes finely areolate. Cephalic dorsum areolate, with costae over the ground sculpture, which become rugose on the posterior sixth of the head; concentric rugulae surrounding the antennal insertions. Lateral surfaces of head areolate, with coarse rugae over the ground sculpture. Ventral surface of head smooth and shining medially, but otherwise weakly areolate with weak costulae. Pronotal neck areolate. Anterior face of pronotum weakly areolate. Lateral surfaces of the mesosoma sculptured similarly to the dorsum of the head. Propodeal declivity areolate-strigulate. Mesoscutum and mesoscutellum with fine costulae over finely areolate ground sculpture. Metanotum finely areolate. Propodeum areolate, with coarse costae. Femora shining through weak areolate sculpture. Petiole uniformly areolate; a weak carina present laterally, extending longitudinally from the petiolar spiracle to the caudal cylinder; coarse rugae on lateral faces of the node. Postpetiole uniformly areolate, with weak rugose sculpture on the dorsal and lateral faces. First gastral tergite shining through weak areolate sculpture; without spectral iridescence. First gastral sternite smooth and shining.

Setae: antennal scapes and funiculi with short, subdecumbent pilosity. Dorsum of the head, pronotum, waist segments, and gaster with moderately abundant, erect, blunt-tipped setae, the longest of which are about a third of the width of the compound eye. Short, sparse pubescence present over the entire body, but difficult to detect against the densely sculptured integument.

Color: predominantly testaceous yellow, with legs lighter yellow. Wing bases dark brown.

Etymology: Morphological, from the Latin ‘rugosus’ (= wrinkled), in reference to the rugose head and mesosoma ( Mackay, 2000).

Comments: From Mackay (2000): “The type series was collected in a trap baited with Vienna sausage, placed in oak trees, about 2-meter height. The habitat was a very steep, south-facing slope of a mountain, covered with oak trees. The specimens were collected within 20 meters distant from one another.” After inspecting the collections of the LACM, I found that W.S. Creighton collected Temnothorax rugosus in a number of localities spanning northern Mexico and southwestern Texas, in addition to the type locality near Monterrey. Like members of the annexus group, which it is closely affiliated with, T. rugosus nests arboreally in the branches a number of different Quercus spp. (including Q. canbyi , Q. clivicola , Q. grisea , and Q. polymorpha ), as well as Cercis reniformis and Rhus spp. at mid elevations. Temnothorax rugosus is also recorded as being collected from Tillandsia carlos-hankii in oak-pine forest near Ixtepeji, Oaxaca ( Franco Méndez, 2008). The femora of Temnothorax rugosus , as noted above, tend to be incrassate, but the degree of swollenness appears to scale with the size of the worker, with smaller workers having less incrassate femora. Temnothorax rugosus shows some variability in petiolar node shape, eye size, and sculpture among populations ( Figs. 152G–152L ). In the petiolar node, there appears to be a complete range of variation between completely cuneiform, with no differentiation between the dorsal and posterior declivitous face, to subcuneiform, with a very short but distinct posterior face and a deeply sloping dorsal face.

Figures

Figure 150 Geographical and elevational distribution of the rugosus group. (A) Temnothorax parralensis sp. nov. (B) T. rugosus. Colored scale to the left of each map represents elevation in meters. Points in black represent the species named in each subfigure, while points in red represent other members of the species group. Type localities are represented by triangles, non-type localities are represented by circles. Bounding box in inset map shows location of main map. Full-size DOI: 10.7717/peerj.11514/fig-150

Figure 152 Temnothorax rugosus. (A–C) Worker (LACMENT323296). (A) Full-face view. (B) Profile view. (C) Dorsal view. (D–F) Gyne (LACMENT323337). (D) Full-face view. (E) Profile view. (F) Dorsal view. Morphological variant, Texas, U.S.A. (G–I) worker (LACMENT323260). (G) Full-face view. (H) Profile view. (I) Dorsal view. (J–L) Gyne (LACMENT323259). (J) Full-face view. (K) Profile view. (L) Dorsal view. Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-152