Alpiscorpius gamma ( Di Caporiacco, 1950 ) Kovařík & Štundlová & Fet & Šťáhlavský, 2019

Kovařík, František, Štundlová, Jana, Fet, Victor & Šťáhlavský, František, 2019, Seven new Alpine species of the genus Alpiscorpius Gantenbein et al., 1999, stat. n. (Scorpiones: Euscorpiidae), Euscorpius 287, pp. 1-29 : 16

Alpiscorpius gamma ( Di Caporiacco, 1950) , comb. n.

( Table 1) http://zoobank.org/urn:lsid:zoobank.org:act:BFD7E5B7-

981B-48DC-8827-559563CE14B4

? Euscorpius germanus “forma C”: Capra, 1939: 204 (part: Tarnova della Selva; now Slovenia, Trnovo).

Euscorpius germanus gamma Di Caporiacco, 1950: 214 .

Euscorpius mingrelicus gamma: Fet & Sissom, 2000: 370 .

Euscorpius gamma: Scherabon et al., 2000: 253 ( Slovenia, in part); Fet et al., 2001: 264 ( Slovenia, in part); Fet et al., 2004: 55 ( Slovenia, in part).

= Euscorpius mingrelicus caprai Bonacina, 1980: 73–75 (see Fet & Sissom, 2000: 370). http: //zoobank. org/ urn : lsid: zoobank. org: act: F39631A9-A5E3-4B71- 9D26-7C85808D084F

TYPE LOCALITY AND TYPE DEPOSITORY. Slovenia, Goriška : north Istria, mouth of the Rižana (Risano) River ; MZUF.

TYPE MATERIAL. Lectotype ♀ (designated by Scherabon et al., 2000: 259) ( MZUF 5581 ), Slovenia: Goriška , north Istria , mouth of the Rižana River . Paralectotypes (11 specimens, MZUF): Slovenia (mouth of the Rižana River , Kamno, Vrsno, Krn, Trenta Valley , and Panovizza, now Panovec), Italy (Udine: Cave di Predil ), and unspecified localities in Soča (Isonzo) River Valley in Slovenia [possibly A. sigma sp. n.] .

DISTRIBUTION. Slovenia (Goriška);? Italy,? Croatia.

NOTES. The karyotypic data for this species have not been published by ŠtundlovÁ et al. (2019). Currently, we restrict Alpiscorpius gamma (Di Caporiacco) only to its type locality in southwestern Slovenia, close to the Croatian border. According to our unpublished data, populations from this area has 2n = 78; it is difficult at this moment to further characterize A. gamma s. str. due to the lack of material. The paralectotype series likely includes members of other cryptic species; the specimens reported from NE Italy (Udine; see also Vignoli & Salomone, 2008, fig. 27) possibly refer to A. sigma sp. n. (so far the only confirmed “ gamma group” species in Italy; its holotype originates from Studena Bassa in Udine). The same may be true for a population from Tarnova della Selva (now Slovenia, Trnovo) mentioned by Capra (1939) and Bonacina (1980).

Fet et al. (2001) listed numerous Slovenian populations of “ Euscorpius gamma ” and noted their “patchy distribution” across the compleX mountainous landscapes. These records, as we see now, could refer to at least five cryptic, allopatric species of the “ gamma group”, four of which are described below.

In the western part of Slovenia, Alpiscorpius species of the “ gamma group” (probably A. omikron sp. n.) were found to be sympatric or maybe even syntopic with species of the “ germanus group” ( A. kappa sp. n. and A. lambda sp. n.).