Macrocheilus tripustulatus (Dejean, 1825)

Zhao, Danyang & Tian, Mingyi, 2012, Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines, ZooKeys 228, pp. 77-100 : 86-87

Macrocheilus tripustulatus (Dejean, 1825) Figs 6, 14, 22, 31, 32, 41, 52

Helluo tripustulatus Dejean 1825: 286 (syn. in Reiche 1842: 334). Reiche 1842: 334; Chaudoir 1872: 212; Heller 1900: 3; Andrewes 1919: 124; Andrewes 1920: 501; Csiki 1932: 1575; Lorenz 2005: 512. Type locality: Indonesia (Java), deposited in MNHN.

Diagnosis.

Length 12.0-12.5 mm, width 4.0-4.3 mm. Labrum (Fig. 6) strongly convex on anterior part, rounded and pointed in front, front seta on apical margin, intermediate one on upper surface close to margin; mandibles sharp at apex; mentum (Fig. 14) irregular and densely setose, both tooth and lobes stout, tooth shorter than lobes, densely setose at basal half; palpi slender, palpomere 4 (Fig. 22) not dilated, densely setose, rounded and truncate at apex, labial palpomere 3 not dilated, bisetose inside; ligula strongly thickened and dilated, deeply depressed at sides, apex rounded, slightly emarginate in middle with a seta on either side. Elytra with two pairs of spots, front spot rounded, covering intervals 3-7 in the middle, hind spot close to apical inner angles, nearly rectangular, covering intervals 1-5; apex roundly truncate.

Male genitalia. Median lobe dilated in the middle ventrally, apical lamella long, wide and round at apex (Figs 31-32).

Female genitalia: Gonocoxite slender, setose on ventral surface, three setae on dorsal side, apex sharp (Fig. 41).

Remarks.

Macrocheilus tripustulatus can be distinguished from Macrocheilus parvimaculatus by the slender maxillary palpomere 4, greater convexity of labrum and larger elytral spots.

Materials examined.

1 female, the holotype, "tripustulatus Dejean (non Fabr.), Java, Coll. Dejean, 3-pustulatus Wiedemann ( Helluo ). Fabr. (Brachinus), Western, Ex-Musaeo Chaudoir"; 1 male, "Java merid., 1500, 1891, H. Fruhstorfer."; 1 male, " Java", "Ex-Musaeo Mniszech"; 1 female, "Carin Chebà, 900-1100 m, L. Fea, V Xii-88; Ex-Musaeo H.W. Bates 1892 ". All specimens deposited in MNHN.

Distribution.

Myanmar and Indonesia (Java).