Exocelina evelyncheesmanae Shaverdo, Hendrich & Balke

Shaverdo, Helena V., Surbakti, Suriani, Hendrich, Lars & Balke, Michael, 2012, Introduction of the Exocelina ekari-group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae), ZooKeys 250, pp. 1-76 : 16-18

12. Exocelina evelyncheesmanae Shaverdo, Hendrich & Balke sp. n. Figs 3 A–E, 29

Type locality.

Indonesia: West Papua Province: Raja Ampat Regency, Waigeo Island, Mountain Nok.

Type material.

Holotype: male "N.DUTCH NEW GUINEA: Waigeu.Camp 1.Mt.Nok. 2,500 ft. v.1938. L.E.Cheesman. B.M.1938-593." (BMNH). Paratypes: 19 males, 16 females with the same labels as the holotype (BMNH, NHMW). 3 males, 2 females "N.DUTCH NEW GUINEA: Waigeu. Mt.Nok. Camp 2. (Buffelhorn.)vi.1938. L.E.Cheesman. B.M.1938-593." (BMNH). 22 males, 24 females "N.DUTCH NEW GUINEA: Waigeu. Camp Nok. 2,500 ft. iv.1938. L.E.Cheesman. B.M.1938-593." (BMNH, NHMW). 18 males, 44 females "Indonesia: Papua, Waigeo, Waifoi, Mt. Nok, 500m, 11.ii.1994, 00.05.076S, 130.44.586E, Balke (BH 11)", one of them additionally with a green label "DNA M.Balke 1276" (NHMW, ZSM).

Diagnosis.

Beetle middle-sized, dark brown to piceous (teneral specimens reddish-brown), shiny; pronotum with lateral bead; male antennomeres 3-7 very slightly enlarged, antennomere 3 slightly more triangular than other antennomeres; male protarsomere 4 with middle-sized, slender, evidently curved anterolateral hook; median lobe with strong submedian constriction in ventral view and truncate apex in lateral view; paramere with notch on dorsal side and subdistal part short and small, with less numerous, relatively short, thick, and flattened setae. The species occurs together with Exocelina waigeoensis sp. n. and can be distinguished from it by its larger size (also females) and the shape of the medial lobe.

Description.

Size and shape: Beetle middle-sized (TL-H 3.75-4.1 mm, TL 4.05-4.45 mm, MW 1.9-2.2 mm), with oblong-oval habitus, broadest at elytral middle. Coloration: Head reddish-brown, darker medially and posterior eyes, to piceous, with dark brown anterior margin, pronotum reddish-brown to piceous, paler on sides (in teneral specimens anterior angles yellowish-red), elytra dark brown to piceous, seldom with reddish sutural bands, head appendages yellow to yellowish-red, legs distally darker than head appendages, hind legs dark brown (Fig. 29).

Surface sculpture: Head with dense punctation (spaces between punctures 1-4 times size of punctures), evidently finer and sparser anteriorly; diameter of punctures smaller than diameter of cells of microreticulation. Pronotum with finer, sparser, and more evenly distributed punctation than on head. Elytra with very sparse and extremely fine punctation. Pronotum and elytra with weakly impressed microreticulation, dorsal surface, thus, shiny. Head with microreticulation stronger. Metaventrite and metacoxa distinctly microreticulate, metacoxal plates with longitudinal strioles and transverse wrinkles. Abdominal sternites with distinct microreticulation, strioles, and fine sparse punctation, coarser and denser on two last abdominal sternites.

Structures: Pronotum with distinct lateral bead, absent in anterior angles. Base of prosternum and neck of prosternal process with distinct ridge, smooth and slightly rounded anteriorly (less than in Exocelina waigeoensis sp. n.), without anterolateral extensions. Blade of prosternal process lanceolate, relatively narrow, convex, with distinct bead and few setae; neck and blade of prosternal process evenly jointed. Abdominal sternite 7 broadly rounded apically.

Male: Antennomeres 3-7 very slightly enlarged, antennomere 3 slightly more triangular than other antennomeres (Fig. 3A); antennomeres 3-5 rugose ventrally. Protarsomere 4 with middle-sized, slender, evidently curved anterolateral hook. Protarsomere 5 ventrally with anterior row of 13 short setae and posterior row of 5-6 short setae (Fig. 3B). Abdominal sternite 7 with 5-12 lateral striae on each side. Median lobe with strong submedian constriction in ventral view and truncate apex in lateral view (Figs 3C, D). Paramere with notch on dorsal side and subdistal part short and small, with less numerous, relatively short, thick, and flattened setae (Fig. 3E).

Female: Antennae more slender, abdominal sternite 7 without striae.

Distribution.

Indonesia: West Papua Province: Raja Ampat Regency. The species is known only from the type locality (Fig. 50).

Etymology.

The species is named for the incredible collector and adventurer, Miss Lucy Evelyn Cheesman who discovered this species. The species name is a noun in the genitive case.