Codorcas antanosy, Barbosa, Diego N. & Azevedo, Celso O., 2012

Barbosa, Diego N. & Azevedo, Celso O., 2012, Revision of Mesitiinae (Hymenoptera, Bethylidae) from Madagascar, with description of eleven new species, Zootaxa 3417, pp. 1-44 : 2

Key to Mesitiinae species from Madagascar

Males (unknown for Codorcas antanosy sp. nov. and Pycnomesitius tanala sp. nov.)

1. Hypopygial median stalk undivided ( Figs 57–62 ). (Tribe Mesitini).............................................. 2

- Hypopygial median stalk partial divided ( Fig. 64–66 ). (Tribe Domonkosini)....................................... 8

2. Sublateral carina of propodeal disc absent ( Figs 12, 15 , 20, 23 ). ( Clytrovorus )...................................... 4

- Sublateral carina of propodeal disc present................................................................. 3

3. Flagellomere 1 longer than pedicel ( Figs 30 , 35 ). (Itapayos).................................................... 7

- Flagellomere 1 shorter than pedicel ( Figs 3, 6 ). ( Anaylax )...................................................... 6

4. Propodeum with posterior spine.......................................................................... 5

- Propodeum without posterior spine ( Fig. 24 )............................................... C. zafimaniry sp. nov.

5. Body castaneous; head with frontal carina ( Figs 17 , 19–21 )...................................... C. merina sp. nov.

- Body dark castaneous nearly black; head without frontal carina ( Figs 9–16 )........................... C. bara sp. nov.

6. Macropterous ( Fig. 1 ); fore wing without costal vein........................................... A. betsileo sp. nov.

- Brachypterous ( Fig. 2 ).................................................................. A. mahafaly sp. nov.

7. Macropterous ( Fig. 26 ) propodeal disc without longitudinal ridge between median and inner discal carina ( Fig. 31 )............................................................................................. I. antaimoro sp. nov.

- Micropterous ( Fig. 33 ); propodeal disc with longitudinal ridge between median and inner discal carina ( Fig. 36 )................................................................................................... I. mikea sp. nov.

8. Metasomal tergum 2 densely granulated ( Fig. 51 ).............................. Pilomesitius madagascarensis Móczár

- Metasomal tergum 2 sparsely punctured ( Figs 41, 42 , 53 ) (Zimankos)............................................ 9

9. Flagellomere 1 at least 2.0 × longer than others ( Fig. 43 ); mesoscutum without longitudinal mesoscutal furrow ( Fig. 44 )........................................................................................... Z. makoa sp. nov.

- Flagellomere 1 at most 1.2 × longer than others; mesoscutum with longitudinal mesoscutal furrow ( Figs 47 , 55 )......... 10

10. Median carina of propodeal disc complete ( Fig. 55 ); hind wing with five distal hamuli....... Z. madagascarensis (Móczár)

- Median carina of propodeal disc incomplete ( Fig. 47 ); hind wing with three distal hamuli............. Z. sakalava sp. nov.

Females (unknown for Anaylax betsileo sp. nov., A. mahafaly sp. nov., Clytrovorus merina sp. nov., C. zafimaniry sp. nov., Itapayaos antaimoro sp. nov., I. mikea sp. nov., Zimankos makoa sp. nov., Z. sakalava sp. nov.)

1. Metasomal tergum 2 densely sculptured ( Figs 34 , 52 )......................................................... 2

- Metasomal tergum 2 not sculptured ( Figs 10 , 25 )............................................................. 3

2. Macropterous; metasomal tergum 2 densely punctured ( Fig. 34 )......................... Pycnomesitius tanala sp. nov.

- Brachypterous ( Fig. 49 ); metasomal tergum 2 densely lineated and hairy ( Fig. 51 ).... Pilomesitius madagascarensis Móczár

3. Propodeal disc without inner discal carina ( Fig. 28 )...................................... Codorcas antanosy sp. nov.

- Propodeal disc with inner discal carina ( Fig. 15 )............................................................. 4

4. Propodeal disc without sublateral and lateral carinae ( Fig. 15 ).............................. Clytrovorus bara sp. nov.

- Propodeal disc with sublateral and lateral carinae............................... Zimankos madagascarensis (Móczár)

Figures

FIGURES 57 – 66. Hypopygium. 57. Anaylax mahafaly sp. nov.; 58. Clytrovorus bara sp. nov.; 59. C. merina sp. nov.; 60. C. zafimaniry sp. nov.; 61. Itapayos antaimoro sp. nov.; 62. I. mikea sp. nov.; 63. Zimankos makoa sp. nov.; 64. Z. sakalava sp. nov.; 65. Pilomesitius madagascarensis Móczár; 66. Z. madagascarensis (Móczár). Scale bars = 300 µm.

FIGURES 11 – 16. Clytrovorus bara sp. nov. 11 – 13. Holotype, 3. 11. Head in dorsal view; 12. Mesosoma in dorsal view; 13. Mesosoma in lateral view; 14 – 16. Paratype, Ƥ. 14. Head in dorsal view; 15 Mesosoma in dorsal view; 16. Mesosoma in lateral view. Scale bars = 300 µm.

FIGURES 19 – 24. 19 – 21. Clytrovorus merina sp. nov., 3, holotype. 19. Head in dorsal view; 20. Mesosoma in dorsal view; 21. Mesosoma in lateral view; 22 – 24. C. zafimaniry sp. nov., 3, holotype. 22. Head in dorsal view; 23. Mesosoma in dorsal view; 24. Mesosoma in lateral view. Scale bars = 300 µm.

FIGURES 27 – 32. 27 – 29. Codorcas antanosy sp. nov., Ƥ, holotype. 27. Head in dorsal view; 28. Mesosoma in dorsal view; 29. Mesosoma in lateral view; 30 – 32. Itapayos antaimoro sp. nov., 3, holotype. 30. Head in dorsal view; 31. Mesosoma in dorsal view; 32. Mesosoma in lateral view. Scale bars = 300 µm.

FIGURES 35 – 40. 35 – 37. Itapayos mikea sp. nov., 3, holotype. 35. Head in dorsal view; 36. Mesosoma in dorsal view; 37. Mesosoma in lateral view; 38 – 40. Pycnomesitius tanala sp. nov., Ƥ, holotype. 38. Head in dorsal view; 39. Mesosoma in dorsal view; 40. Mesosoma in lateral view. Scale bars = 300 µm.

FIGURES 3 – 8. 3 – 5. Anaylax betsileo sp. nov., 3, holotype. 3. Head in dorsal view; 4. Mesosoma in dorsal view; 5. Mesosoma in lateral view; 6 – 8. A. mahafaly sp. nov., 3, holotype. 6. Head in dorsal view; 7. Mesosoma in dorsal view; 8. Mesosoma in lateral view. Scale bars = 300 µm.

FIGURES 17 – 18. 17. Clytrovorus merina sp. nov., 3, holotype, habitus; 18. C. zafimaniry sp. nov., 3, holotype, habitus. Scale bars = 300 µm.

FIGURES 9, 10. Clytrovorus bara sp. nov., habitus (9. 3, holotype; 10. Ƥ, paratype). Scale bars = 300 µm.

FIGURES 1 – 2. 1. Anaylax betsileo sp. nov., 3, holotype, habitus; 2. Anaylax mahafaly sp. nov., 3, holotype, habitus. Scale bars = 300 µm.

FIGURES 25 – 26. 25. Codorcas antanosy sp. nov., Ƥ, holotype, habitus; 26. Itapayos antaimoro sp. nov., 3, holotype, habitus. Scale bars = 300 µm.

FIGURES 33 – 34. 33. Itapayos mikea sp. nov., 3, holotype, habitus; 34. Pycnomesitius tanala sp. nov., Ƥ, holotype, habitus. Scale bars = 300 µm.

FIGURES 51 – 52. 51. Pilomesitius madagascarensis Móczár (51. Ƥ, holotype, metasoma dorsal; 52. 3, paratype, metasoma dorsal). Scale bars = 300 µm.

FIGURES 41 – 42. 41. Zimankos makoa sp. nov., 3, holotype, habitus; 42. Z. sakalava sp. nov., 3, holotype, habitus. Scale bars = 300 µm.

FIGURES 53 – 56. Zimankos madagascarensis (Móczár), 3 53. Habitus; 54. Head in dorsal view; 55. Mesosoma in dorsal view; 56. Mesosoma in lateral view. Scale bars = 300 µm.

FIGURES 43 – 48. 43 – 45. Zimankos makoa sp. nov., 3, holotype. 43. Head in dorsal view; 44. Mesosoma in dorsal view; 45. Mesosoma in lateral view; 46 – 48. Z. sakalava sp. nov. 3, holotype. 46. Head in dorsal view; 47. Mesosoma in dorsal view; 48. Mesosoma in lateral view. Scale bars = 300 µm.

FIGURES 49 – 50. 49. Pilomesitius madagascarensis Móczár, female holotype, habitus; 50. Pilomesitius madagascarensis Móczár, male paratype, habitus. Scale bars = 300 µm.