Pupina exclamationis Mabille, 1887

Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak, 2022, Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia, ZooKeys 1119, pp. 1-115 : 1

Pupina exclamationis Mabille, 1887

Figs 29I-K , 34B, C

Pupina exclamationis Mabille, 1887: 137, 138, pl. 4, figs 11, 12. Type locality: Tonkin. Fischer 1891: 108. Fischer and Dautzenberg 1904: 431, Bac-Kan, Tonkin; Monts Mauson, Tonkin [Mount Mau Son, Lang Son Province, Vietnam]. Do et al. 2015: 126, fig. 6a, Son La Province, Vietnam.

Pupina (Tylotoechus) exclamationis - Kobelt 1902: 312.

Type material examined.

Syntypes MNHN-IM-2000-35840 (4 shells; Figs 29I, J , 34B ) from Tonkin.

Other material examined.

NHMUK 1901.12.23.205-210 "forma minor" ex. H. Fruhstorfer coll. (5 shells; Figs 29K , 34C ) from Than-Moi, Tonkin.

Diagnosis.

Shell ovate-fusiform to fusiform; last whorl ca. three quarters of shell height. Apertural lip somewhat thickened but not expanded; apertural lip slightly curved when observed from lateral view. Columellar tooth fin-shaped.

Differential diagnosis.

Pupina exclamationis is most similar to P. peguensis in having a glossy surface and a curved apertural lip when observed from lateral view, but differs in having a more fusiform shell shape and a less distinct parietal callus.

Distribution.

Northern Vietnam ( Do et al. 2015).

Figures

Figure 29. Shells of Pupina arula species group from mainland Southeast Asia A, B Pupina arula A syntype UMZC I. 103025 and B specimen NHMUK 1888.12.4.109. C, D Pupina mouhoti, possible syntypes NHMUK ex. Cuming coll. E, F Pupina vescoi E syntype NHMUK 1893.2.4.767 and F specimen SMF 109956 / 1 G Pupina perakensis, lectotype SMF 109969 / 1 H Pupina excisa, lectotype SMF 110778 / 1 I-K Pupina exclamationis I, J syntypes MNHN-IM- 2000 - 35840 and K specimen NHMUK 1901.12.23.205 " forma minor ". Photo: J. Ablett, H. Taylor, NHM (A), A. Lardeur, P. Maestrati, MNHN (I, J).

Figure 34. A Pupina vescoi, specimen SMF 109956 / 1 from Cochin China B, C Pupina exclamationis B syntype MNHN-IM- 2000 - 35840 from Tonkin and C specimen NHMUK 1901.12.23.205 " forma minor " from Than-Moi, Tonkin D Pupina perakensis, lectotype SMF 109969 / 1 from Bukit Pondong, Perak E Pupina excisa, lectotype SMF 110778 / 1 from Kelantan F Pupina aureola, possible syntype NHMUK 1988.12.4.101 from Pinang. Photo: P. Maestrati, MNHN (B).