Glypturus acanthochirus Stimpson, 1866

Klompmaker, A., Hy ̆ z, Matu ̆ s, , Roger W. Port & Kowalew, nd Michał, 2016, Growth, inter- and intraspecific variation, palaeobiogeography, taphonomy and systematics of the Cenozoic ghost shrimp Glypturus, Journal of Systematic Palaeontology 14 (2), pp. 99-126 : 114-116

Glypturus acanthochirus Stimpson, 1866

( Fig. 15)

‡ 1866 Glypturus acanthochirus Stimpson , 46.

2012 Glypturus acanthochirus Stimpson ; Hy̆zńy & Muller: 971, figs 1, 2A — C, 3A — C, I, 4C (cum syn.).

Diagnosis. Front trispinous. Manus of large cheliped usually with three well-developed spines along distal half of dorsal margin. Endopod of uropod elongate oval, about twice as long as wide. Outer side major propodus with tubercles arranged on lower half (~8 — 17% of manus), not reaching proximal corner. Tubercles on inner side major propodus absent to rare (<~6% manus). Carpi without tubercles on outer side; part of specimens exhibit tubercles near lower-distal margin on inner side (<~5% coverage). Minor chelipeds without tubercles (expanded from Biffar 1971, p. 655).

Material. F = fossil (propodi), remainder = modern (entire specimens). NHMW 6766 (Twin Cays, Belize), 6768 — 6770 (Twin Cays, Belize), 15338 (Twin Cays, Belize), 15340 ( Fig. 15A — D; Twin Cays, Belize), 15341 — 15342 (Twin Cays, Belize), 19624 (Twin Cays, Belize), 19625 (South Water Caye, Belize), 19626 — 19627 (Twin Cays, Belize), 24967 ( Fig. 15E, F; Twin Cays, Belize), 24968 (Twin Cays, Belize), 25261 (Pos Chiquito, Aruba), 25266 (Cayo Muerto, Morrocoy, Venezuela), 25635 (near STRI station, Bocas del Toro, Panama), 25636 ( Panama, Bocas del Toro), 25637 ( Panama, Bocas del Toro, San Cristobal, Punta Coco), 25638 ( Panama, Bocas del Toro), 25639 — 25640 ( Panama, Isla Grande). MNHN Th-1593 ( Venezuela, Isla Margarita, Guamache). UF 32050 (French Antilles, Saint Martin, Le Galion), 32479 (French Antilles, Saint Martin, Little Key), 32113 (French Antilles, Saint Martin, Le Galion), 32131 (French Antilles, Saint Martin, Baie de l’Embrouchure), 32483 (French Antilles, Saint Martin, Little Key), 42451 (F, Fig. 15G — I; upper Pliocene — lower Pleistocene Tamiami Formation, Pinecrest beds, of Florida, Mule Pen), 232383 (F, Fig. 15J — M; Holo-Pleistocene of Jamaica, Falmouth 01), 232384 — 232385 (two fixed fingers, Holo-Pleistocene, Jamaica, Falmouth 01).

Occurrence. Western Atlantic region: Gulf of Mexico, Caribbean region, and northern part of the east coast of South America ( Dworschak 1992; Sakai 2005; Hy̆zńy & Muller 2012). Fossil specimens known from the Pleistocene and Holo-Pleistocene of Jamaica, and the late Pliocene — early Pleistocene of Florida ( Collins et al. 1996; herein) (Supplemental Table 1 ).

Description. Additional to Biffar (1971, pp. 655 — 660): major propodus exhibiting tooth on fixed finger; tooth on fixed finger of minor reduced to absent. Upper margin minor and major propodus exhibit usually three forwardly oriented spines. Outer side major propodus with tubercles arranged on lower half (~8 — 17% of manus), not reaching proximal corner. Tubercles on inner side major propodus absent to rare (<~6% manus). Carpi without tubercles on outer side; part specimens exhibit tubercles near lower-distal margin on inner side (<~5% coverage). Meri with tubercles below longitudinal keel (especially proximally) on outer side of major propodus for larger specimens; inner side exhibiting tubercles, especially proximally and on lower margin inner plate for larger specimens. Minor chelipeds without tubercles. Fixed finger of minor comprises a statistically greater portion of propodal length compared to fixed finger of major ( Fig. 11A).

Remarks. Biffar (1971, pp. 655 — 660) provided a detailed description of the entire body of the modern species. Details of tuberculation patterns, known to be important for fossil Glypturus spp. , were not described in detail. Therefore, the description of the species is augmented.