Orthogonius carinatus Tian & Deuve

Tian, Mingyi & Deuve, Thierry, 2013, Definition and review of the lancangjiang species group of the termitophilous genus Orthogonius Macleay, 1825 (Coleoptera, Carabidae, Orthogoniini), ZooKeys 349, pp. 81-100 : 89-93

Orthogonius carinatus Tian & Deuve sp. n. Figs 1, 8-9, 18-19

Diagnosis.

Medium to large sized, body elongate, whole dorsal surface extraordinarily and densely punctate; head as long as wide, eyes moderate, labrum roundly protruding at front; pronotum narrow, lateral expanded margins distinctly reflexed throughout; elytra carinate on basal 1/3 of interval 7 in both sexes, and at base of intervals 4-6 in female, but normal in male.

Length: 14.0-16 mm; width: 5.5-6.0 mm. Body slender and elongate. Habitus as in Figs 8-9.

Description.

Black, antennomeres 1, 3-4, legs (except tarsomeres) and ventral surface dark brown, mandibles, labrum, palps, other antennomeres and tarsomeres reddish brown.

Macrosculpture: Head extraordinarily and densely punctate, pronotum distinctly punctate in male, but faintly punctate or impunctate in female; elytra with tiny, sparse and faint punctures in both sexes; underside smooth and glabrous.

Microsculpture: Engraved meshes densely isodiametric or slightly transverse on elytra, faint or not clear on head and pronotum.

Head stout, as long as wide,eyes moderate, prominent, frons and vertex moderately convex, frontal impressions not well defined, shallow; clypeus bisetose, base area rugose; labrum broadly protruding at front, sexsetose; mandibles rather short; ligula broad, bisetose at apex; maxillary palps moderate, subcylindrical, maxillary palpomere 3 slightly longer than 4, both glabrous except for palpomere 3 with two short setae at apex; labial palpomeres stout, palpomere 3 somewhat expanded, palpomere 2 cylindrical, as long as palpomere 3, bisetose on inner margin, and with two or three additional setae at subapex and apex; palpiger asetose, mentum and submentum each with one pair of setae; mentum without median tooth. Antennae long and slender, extending beyond basal 1/3 of elytra; pubescent from basal 1/3 of antennomere 4, antennomere 3 slightly shorter than 4.

Pronotum transverse, PW/PL=1.55-1.60, disc moderately convex; sides evenly rounded, widest at middle, both basal and fore margins beaded, lateral expanded margins well defined, wide and obviously reflexed throughout; fore and hind angles rounded; both fore and hind transversal impressions faint, basal foveae large and wide, connected to expanded margins; median line clear.

Elytra elongate, EL/EW=1.72-1.78, moderately convex, base completely bordered; sides nearly parallel at middle, striae deep, intervals convex, subequal in width in middle; apex broadly truncate; carinate on bases of intervals 6-8, basal 1/3 of interval 7 in both sexes, and at base of intervals 4-6 in female (carination shortest on interval 4, and longest on 6); interval 3 with only the basal and middle setiferous pores, the subapical one absent.

Legs slender; middle and hind coxae glabrous in median portion; fore tibia slightly dilated at apex, outer angle obtuse, apex slightly and obliquely truncate, outer margin slightly subserrate; middle and hind tibiae hardly dilated at apex, slender and strongly curved in middle portion in male, normal in female; hind tarsomeres 1 and 3 much longer than 2 and 4 respectively, length ratio of tarsomeres 1-4 as 3.9: 2.6: 1.8: 1.0; tarsomere 4 deeply emarginate at apex, lobes asymmetric, longer lobe as long as 1/3 of the joint; all tarsal claws strongly pectinate.

Prosternal process faintly bordered at apex. Abdominal ventrite VII distinctly emarginate at apical margin in male.

Male genitalia (Figs 18-19). Rather slender, strongly sinuate ventrally, distinctly arcuate at apical 1/3, slightly dilated in middle portion, dorsal opening large, subapex nearly paralleled towards apex, broadly obtused at apex; in dorsal view, apical lamella almost as long as wide, not parallel–sided, apex broadly rounded.

Remarks.

Similar to Orthogonius lancangjiang , but intervals of elytra carinate; the median lobe of aedeagus distinctly arcuate and the apical lamella somewhat stouter. It is easily distinguished from the next species, Orthogonius macrophthalmus sp. n., by its small and less prominent eyes, narrower pronotum, and elytra carinate on base of intervals 4-6 in female.

Material examined.

Holotype: male, labeled: “Laos–NE: Houa Phan Province, Ban Saluei to Phou Pane Mt., 20.12 N/ 104.01 E, 1500-1900 m, 17.V.-3.VI. 2007, M. Brancucci Leg.", and "NHMB Basel Expeditionto Laos, 2007", in NHMB.

Paratypes. 1 male and 3 female, idem; 1 male 3 females, “Laos–NE: Houa Phan Province, Ban Saluei to Phou Pane Mt., 20.1309N/ 103.5954-104.0003E, 1480-1550 m, 9-16. VI. 2009, David Hauck Leg.", and "NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kuban"; 1 male and 5 females, “Laos–NE: Xieng Khouang Province, 30 km NE Phonsaven, Phou Sane Mt., 19.37-8N/ 103.20-21E, 1420 m, 30 km NE Phonsaven, Ban Na Lam to Phou Sane Mt., 1300-1700 m, 10.-30.V. 2009, M. Geiser Leg.", and "NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kuban"; 1 male and 2 females, “Laos–NE: Houa Phan Province, Ban Saluei to Phou Pane Mt., 20.13N/ 104.00E, 1350-1500 m, 1-16. VI. 2009, M. Brancucci Leg.", and "NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kuban"; 3 females, “Laos–NE: Xieng Khouang Province, 30 km NE Phonsaven: Ban Na Lam to Phou Sane Mt., 19.37-38N/ 103.20E, 1300-1500 m, 10.-30.V. 2009, M. Brancucci Leg.", and "NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kuban"; 2 female, “Laos–NE: Houa Phan Province, Ban Saluei to Phou Pane Mt., 20.12-13N/ 103.595-104.01E, 1340-1870 m, 10. V.-16. VI. 2009, M. Brancucci and local coll. Leg.", and "NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kuban"; 1 male 6 female, “Laos–NE: Xieng Khouang Province, Phonsaven (30 km NE), Phou Sane Mt., 19.37-38N/ 103.20E, 1400-1500 m, 10.-30.V. 2009, Z. Kraus Leg.", and "NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kuban"; 1 females, “Laos–N: (Oudom Xai), 17 km, NEE of Oudom Xai, 20.45N/ 102.09E, 1-9. V. 2002, V. Kuban leg."; 7 females, “Laos–NE: Xieng Khouang Province, Phonsaven (30 km NE), Phou Sane Mt., 19.37-38N/ 103.20-21E, 1400-1700 m, 10.-30.V. 2009, D. Hauck Leg.", and "NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kuban"; 1 female, “Laos–NE: Houa Phan Province, Ban Saluei to Phou Pane Mt., 20.11-13N/ 103.59-104.01E, 1300-1900 m, 9-17.VI. 2009, Michael Geiser Leg.", and "NHMB Basel, NMPC Prague Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kuban", all in NHMB except two males and ten females in MNHN and SCAU, respectively. 4 female, "Lao, Phongsaly Prov., 21°41-2'N /102°6-8'E, 28. v.-20. vi. 2003, Phongsaly env., 1500 m, Pacholátko leg.", and "Collection Naturhistorisches Museum Basel", all in NHMB except one in MNHN; 1 female, ibid, except “Pacholátko leg" replaced by "Brancucci leg.", in NHMB.

Etymology.

Referring to the character of elytral carination.

Distribution.

Northern Laos (Fig. 1).