Aposphragisma rimba Thoma, 2014

THOMA, MARCO, KRANZ-BALTENSPERGER, YVONNE, KROPF, CHRISTIAN, GRABER, WERNER, NENTWIG, WOLFGANG & FRICK, HOLGER, 2014, The new Southeast Asian goblin spider genus Aposphragisma (Araneae, Oonopidae): diversity and phylogeny, Zootaxa 3798 (1), pp. 1-86 : 59-64

publication ID	https://doi.org/ 10.11646/zootaxa.3798.1.1
publication LSID	lsid:zoobank.org:pub:CB8534BA-89CB-44A6-81E3-3A8927055C7A
DOI	https://doi.org/10.5281/zenodo.4431933
persistent identifier	https://treatment.plazi.org/id/094F426D-FFAC-FFE2-6FC9-F4EE86F8F849
treatment provided by	Plazi
scientific name	Aposphragisma rimba Thoma
status	sp. nov.

Treatment

Aposphragisma rimba Thoma View in CoL , sp. nov.

( Figs. 38–41 View FIGURE 38 View FIGURE 39 View FIGURE 40 View FIGURE 41 )

Type material. HOLOTYPE: male ( PBI_OON 00031408 ), Sarawak ( MALAYSIA), Mulu N.P ., 100 km SEE of Miri, 200 m [04°00′N 114°49′E; prov.], 19.–24.VIII.2003 (Winkler extraction), leg. A. Schulz, deposited at MHNG GoogleMaps . PARATYPES: 2 males ( PBI_OON 00012526 ), collected together with holotype, deposited at NMBE View Materials GoogleMaps ; 1 female ( PBI_OON 00031406 ; abdomen used for examination of internal genitalia), collected together with holotype, deposited at MHNG GoogleMaps ; 4 males & 1 female ( PBI_OON 00031407 ), collected together with holotype, deposited at MHNG GoogleMaps .

Non-type material. 2 males ( PBI_OON 00031404 ; used for SEM), collected together with holotype, deposited at MHNG GoogleMaps ; 2 aberrant females ( PBI_OON0 0031405 ), collected together with holotype, deposited at MHNG GoogleMaps .

Etymology. The species epithet is a noun in apposition meaning ‘pristine forest’ or ‘jungle’ in the Malay language.

Diagnosis. A. rimba sp. nov. does not belong to the stripe clade and thus shows a completely ornamented sternum. It can be distinguished from congeners with a completely ornamented sternum by the combination of the following traits: entire carapace surface smooth ( Fig. 38D View FIGURE 38 ); carapace margin with blunt denticles, sluice reaching from posterior margin to level of coxa I ( Fig. 38D View FIGURE 38 ); epigastric scutum dorsally without a widely oval subterminal ridge ( Fig. 40C View FIGURE 40 ); booklung covers (bc) small ( Figs. 40 View FIGURE 40 C–E).

Description. Description based on 9 males and 4 females.

MALE. With the characters of the genus except as noted. Body length 1.18–1.45 mm (n=9), carapace length 0.59–0.66 mm (n=9). Sclerotized parts uniformly coloured orange-brown close to colour 36 ‘amber’; legs paler than body, yellowish. Males show several remarkable modifications compared to congeners. Habitus: Figs. 38 View FIGURE 38 A–C.

CEPHALOTHORAX. Carapace: entire surface smooth, pars thoracica with 3 pairs of pronounced narrow depressions along margin ( Fig. 38D View FIGURE 38 ); posterolateral surface without spikes; posterior edge of pars cephalica with slightly stronger modified hair bases, appearing as small denticles; carapace margin with narrow sluice reaching from posterior margin to level of coxa I, with blunt denticles ( Fig. 38D View FIGURE 38 ). Eyes: posterior eye-row straight from above, procurved from front; ALE largest, separated by their radius to diameter; ALE-PLE separated by less than ALE radius; PME touching throughout most of their length; PLE-PME separated by less than PME radius. Sternum completely coarsely ornamented ( Figs. 38 View FIGURE 38 E–G); infracoxal grooves (icg) at both ends with depressions (presence or absence of openings can not be verified on available SEM images) ( Fig. 39C View FIGURE 39 ); posterior margin with broad single extension, covered with blunt denticles ( Fig. 38F View FIGURE 38 ); setae darkish, appearing slightly plumose at SEM level ( Fig. 39C View FIGURE 39 ). Pleura (pl): surface smooth with cluster of openings (cpo) ( Figs. 39 View FIGURE 39 C–D). Mouthparts: chelicerae posterior margin of inner surface proximally modified to a ridge with median slit (sli), covering about 1/2 of margin length ( Fig. 39G View FIGURE 39 ); promargin with row of flattened setae (rfs) in combination with plumose setae ( Fig. 39G View FIGURE 39 ), distally extending into a long tooth-like projection (tlp) ( Figs. 38H View FIGURE 38 , 39G View FIGURE 39 ); anterior face with few thickened and darkish needle-like setae ( Figs. 38H View FIGURE 38 , 39A, G View FIGURE 39 ; cf female, fig. 39B); posterior face with two spatulate setae (sps), about as long as fang ( Fig. 39G View FIGURE 39 ). Labium longitudinally slightly compressed and not as long as in females ( Fig. 39E View FIGURE 39 ). Endites distally with brush of special wing-like setae (ws) on inner margin ( Fig. 39F View FIGURE 39 ), outer margin subdistally with a pair of remarkably modified, darkish and thick setae (ebs) which resemble the blossom of a bird of paradise flower ( Strelitziaceae : Strelitzia sp.; Fig. 39F View FIGURE 39 ): a rather long and flat distal part covered with little spines and filaments is subbasally attached in a 90 degree bend to an unmodified but rather thick basal part ( Figs. 39 View FIGURE 39 E–F).

ABDOMEN. Book lung covers (bc) small, ovoid ( Figs. 40C, E View FIGURE 40 ). Epigastric scutum dorsally without subterminal ridge ( Fig. 40C View FIGURE 40 ). Postepigastric scutum almost semicircular, with long posteriorly directed lateral apodemes (lap) ( Fig. 40A View FIGURE 40 ). Setae appearing slightly plumose at SEM level ( Fig. 40D View FIGURE 40 ).

GENITALIA. Epigastric region: sperm pore (sp) situated between anterior and posterior spiracles ( Fig. 40A, D View FIGURE 40 ). Palp: basal segments as well as bulb and cymbium lighter in colour than rest of body; bulb slender with a faint conical bulge (cb) posterior of embolus-conductor-complex (compared to congeners shifted towards prolateral surface; Figs. 41 View FIGURE 41 D–E). On stereo-microscope level embolus-conductor-complex pincer-like in lateral view, dark ( Figs. 41 View FIGURE 41 A–B). Embolus (em) basally broad, subbasally with an angular dent on ventral part (thus proximal part apearing rhomboid in lateral view), medially abruptly narrowed, without dorsal comb ( Fig. 41C View FIGURE 41 ); embolus (em) tip flattened, elongated spatulate, basally with hook ( Fig. 41C View FIGURE 41 ); conductor (con) slender, attached basally to ventral part of embolus ( Figs. 41C, E View FIGURE 41 ).

FEMALE. As in males except as noted. Body length 1.38–1.44 mm (n=3), carapace length 0.65–0.66 mm (n=4). Chelicerae anterior face with scattered needle-like (and not thickend) setae ( Fig. 39B View FIGURE 39 ). Labium elongated triangular, not compressed. Anterolateral margin of outer surface of endites subdistally with a pair of unmodified long, needle-like setae with tips bent inwards.

GENITALIA. Dorsal view ( Figs. 41 View FIGURE 41 F–G): area containing small circular structures laterally framed by short rectangular sclerites (apodemes, lap); receptaculum (re) small, elongated; a transverse sclerite (ts) lies anteriorly to the receptaculum, medially bearing a nail-like structure (na).

Intraspecific variation. Body size variable (see above); colour shows only very slight variation, one female pale; number of denticles on carapace margin variable. Two females (PBI_OON 00031405) show a pair of tubercles (tub) at the ventral base of the pedicel tube and faint ridges on book lung covers (bc) ( Fig. 40E View FIGURE 40 ).

Distribution. NE-Sarawak, MALAYSIA; all specimens collected at type locality ( Fig. 49 View FIGURE 49 ).