Listrodromus simplex HEINRICH , 1934

Riedel, Matthias, 2023, New contribution to the Oriental species of Anisobas WESMAEL and Listrodromus WESMAEL (Hymenoptera, Ichneumonidae, Ichneumoninae), Linzer biologische Beiträge 54 (2), pp. 625-639 : 631-632

Listrodromus simplex HEINRICH, 1934 (figs 18, 30)

M a t e r i a l e x a m i n e d: Indonesia: C Sulawesi, nr Luwuk, Salodik, c 400 m, 1 ♀ 21- 31.x.1989, Mal. trap 14, C. v. Achterberg, RMNH´89 (Leiden).

D e s c r i p t i o n: Body length 6.8 mm. Length of fore wing 4.9 mm. Flagellum short, filiform, with 20 flagellomeres; 1 st flagellomere (without anellus) 2.2× as long as wide and 1.0× as long as 2 nd flagellomere; 2 nd flagellomere 2.1× longer than wide; preapical flagellomere square. Temple slightly and roundly narrowed behind eye. Vertex, occiput and temple almost smooth. Frons, malar space and gena with sparse punctures. Face densely punctate, covered with whitish setae. Clypeus with sparse punctures, apical margin with large median tooth. Mandible with two large teeth, lower tooth smaller and slightly curved inwards. Malar space 1.5× longer than width of mandibular base. Genal carina reaching mandibular base.

Mesosoma shining, with whitish setae. Pronotum largely smooth, with fine setiferous punctures on upper margin and ventrally, with some coarse rugae on posterior margin. Notaulus impressed on frontal fifth of mesoscutum. Mesoscutum with dense punctures in frontal half, almost smooth in posterior third. Mesopleuron densely rugose-punctate; speculum very large and smooth. Metapleuron densely coarsely rugose-punctate but almost smooth ventrally; juxtacoxal carina strong. Scutellum moderately elevated, almost square, with complete lateral carina and sparse fine punctures. Propodeum completely carinate, spiracle oval. Area superomedia partly smooth, area petiolaris with transverse rugae. Area superomedia hexagonal, 1.1× wider than long; anterior transverse carina reaching at 4/10 of its length. Coxae densely punctate; hind coxa without scopa. Hind femur club-shaped, 3.4× longer than wide. Fore and mid claws strongly pectinate; hind claw simple. Areolet pentagonal, frontal distance between veins 2rs-m and 3rs-m 5× their width; vein 2m-cu reaching its middle. Vein 1cu-a slightly antefurcal.

Metasoma amblypygous. Petiolus without latero-median carina. Postpetiolus moderately widened, without latero-median carina, almost smooth with fine longitudinal rugae medially and punctate laterally. Gastrocoelus strongly impressed, with longitudinal ridges. Thyridium almost transverse, c. 0.9× as long as the interval between thyridia. 2 nd tergite 1.4× wider than long; 3 rd tergite 1.6× wider than long. 2 nd and 3 rd tergites coarsely and densely rugose-punctate, with almost smooth apical margins. 4 th tergite with fine sparse punctures; following tergites almost smooth. Ovipositor sheath rather short, slightly reaching behind metasoma apex.

Color: Flagellum reddish-brown; scape reddish, yellow laterally. Head brownish. Palps ochreous. Mandible basally, lateral spots on clypeus apically, wide facial and frontal orbits spot on outer orbit ivory. Mesosoma brown. Tegula brown. Upper margin of pronotum, subtegular ridge and scutellum ivory. Metasoma reddish-brown; petiolus yellow, with brownish apical spot; postpetiolus ivory. Posterolateral spots on 2 nd and 3 rd tergites and apical spots on 4 th to 7 th tergites ivory. Coxae reddish-brown basally, reddish-yellow apically; hind coxa with yellow dorsal spot. Legs otherwise reddish; hind tarsus brown. Wings hyaline; pterostigma brown.

Key for the known ♀♀ of Oriental Listrodromus species (adopted from RIEDEL 2011):

1 Malar space very long, about 2× longer than width of mandibular base (fig.15). 2 nd tergite 1.2-1.3× longer than wide. 1 st and 2 nd tergites with reddish ground color, following tergites ± black (fig. 27). Java, Laos. ............................. L. javae HEINRICH, 1975

- Malar space shorter, not more than 1.6× longer than width of mandibular base (figs 13- 14, 16-18). 2 nd tergite about as long as wide or transverse. 2 nd tergite with black or brown ground colour OR 2 nd and metasoma entirely red (figs 25-26, 28-30). ....................2

2 Tergites red; only 2 nd and 3 rd tergites with weak and small posterolateral ivory spots (fig. 29). Sulawesi. ............................................................................... L. rufiventris nov.sp.

- Tergites black or brown, often with ivory or yellow spots. .................................................3

3 Mesosoma with extended yellow coloration: metapleuron and propodeum with yellowish spots (fig. 20). Flagellum with 18-19 flagellomeres, basal flagellomeres very short, 1 st flagellomere 1.5-1.7× longer than wide. Malaysia (Pahang, Sabah)....................... ........................................................................................... L. flavomaculatus RIEDEL, 2011

- Mesosoma with less extended ivory spots: metapleuron and propodeum black (figs 19, 21-22). Flagellum with 19-23 flagellomeres, 1 st flagellomere at least 1.7× longer than wide, usually longer. ...........................................................................................................4

4 Mesoscutum with two paramedian ivory or yellowish spots which are sometimes confluent apically. Mesopleuron usually with ivory or yellowish spot on epicnemium. 5 th tergite, often also 4 th tergite without ivory coloration (fig. 28). Malaysia (Padang, Sabah). .................................................................................... L. propodealis RIEDEL, 2011

- Mesoscutum and mesopleuron entirely black. 4 th and 5 th tergites with ivory or yellow spots (figs 25-27, 30)...........................................................................................................5

5 Hind femur and tibia red, not darkened. Hind claw simple, not pectinate. Sulawesi .............. ................................................................................................. L. simplex HEINRICH, 1934

- Hind femur and tibia basally and apically ± darkened; hind tibia with pale yellow subbasal band. Hind claw ± pectinate, at least basally. Widespread in Southern and Southeast Asia. ..................................................................... L. crassipes (CAMERON, 1907)