Marshallicona Cerretti & Pape

Cerretti, Pierfilippo, Badano, Davide, Gisondi, Silvia, Giudice, Giuseppe Lo & Pape, Thomas, 2020, The world woodlouse flies (Diptera, Rhinophoridae), ZooKeys 903, pp. 1-130: 1

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Marshallicona Cerretti & Pape

gen. nov.

Marshallicona Cerretti & Pape   gen. nov. Figs 1C View Figure 1 , 15 View Figure 15

Unambiguous character state changes

(Table 1 View Table 1 , Fig. 20 View Figure 20 ). Global apomorphies: none; local apomorphies: 2:1, 9:1, 43:1, 51:1, 52:1, 56:1, 66:0, 68:1.


Head: head higher than long in lateral view. Facial ridge 0.4 times as long as frons. Ocellar setae virtually absent. Frons approx. 0.8 times as wide as compound eye in dorsal view. Median vertical setae strong, subparallel. Six to eight medially crossed frontal setae, slightly reclinate, descending to level of upper margin of scape. Fronto-orbital plate with few short setulae confined to upper half. Four or five proclinate orbital setae (posterior two slightly lateroclinate). One weak upper lateroclinate orbital seta (usually not distinguishable from uppermost frontal setae). Parafacial bare, at its narrowest point approx. as wide as width of postpedicel. Vibrissal angle receding. Vibrissa well developed, arising at level of lower facial margin. Lower facial margin not sunken though not visible in lateral view. Facial ridge concave with decumbent setulae on lower 1/3-2/5. Face slightly concave, antennae not hidden from view in profile. Antenna approx. as long as height of gena. Postpedicel sub-ovoid, approx. 1.5 times as long as pedicel. Arista bottlebrush-like, trichia longer that maximum diameter of arista. Arista thickened in proximal 1/5 or less; second aristomere at most as long as wide. Lunule hidden by inner anterior margins of fronto-orbital plate. Gena, in profile, approx. 1/2 as high as compound eye. Palpus stout, clavate, with a few thin setulae on apical 1/4.

Thorax: prosternum bare. Postpronotum with two setae. One postsutural supra-alar seta (i.e., first and third post sutural supra-alar setae absent). Scutellum with one pair of well-developed basal setae and one pair of strong, horizontal and crossed apical setae; basal setae placed dorsally with respect to apical setae. Anatergite bare. Subscutellum moderately swollen, not fully sclerotised. Metathoracic spiracular lappets virtually absent. Lower calypter distinctly tongue-shaped (ground-plan trait of Rhinophoridae   ) (Fig. 2E View Figure 2 ). Costal sector cs2 setose ventrally. Costal spine not differentiated from general costal setae. Costal sector cs5 longer than costal sector cs2 (Fig. 15E View Figure 15 ). Vein R1 dorsally setose on distal 1/4. Base of R4+5 entirely bare. Bend of vein M1 shallow, well removed from wing margin. Crossvein dm-m forming an acute angle with proximal section of M4. Vein CuA+CuP not reaching wing margin. Preapical anterodorsal seta of fore tibia approx. as long as preapical dorsal seta. Fore tarsus not compressed. Tibiae of mid and hind leg normally developed. Mid tibia with one anterodorsal seta. Hind tibia with 3 dorsal preapical setae.

Abdomen: slightly elongated, virtually without microtomentum and without distinct marginal and discal setae.

Male terminalia (Fig. 15F-H View Figure 15 ): posterior margin of sternite 5 with a deep median notch; lateral lobe rounded posteriorly. Tergite 6 bare, medially divided into two hemitergites; tergite 6 divided from syntergosternite 7+8 by a wide membrane. Connection between sternite 6 and syntergosternite 7+8 on right side membranous. Cerci well developed, not fused medially. Basal 1/3 of cerci convex and covered with short setae; distal 2/3 straight and narrowly digitiform (branches symmetrically diverging and well separated). Surstylus well developed, lobe-like in lateral view; lateral side of surstylus broadly convex at approx. mid length. Surstylus not fused to epandrium. Bacilliform sclerite firmly fused to laterobasal margin of surstylus. Hypandrial arms not fused medially. Connection between phallic guide and pregonite sclerotised. Postgonite without anterior seta. Epiphallus well developed and attached dorsomedially to basiphallus. Extension of dorsal sclerite of distiphallus entirely fused medially into a single sclerite and proximally fused to dorsal sclerite of distiphallus. Median process of ventral sclerotisation of distiphallus present, divided medially into two hemisclerites, which are both proximally fused to ventral plate of distiphallus. Acrophallus simple and scale-like spinules present lateroventrally.


Neotropical - Ecuador.

Type species.

Marshallicona quitu   Cerretti & Pape, sp. nov., by present designation.


The generic name is a composite word formed from the name of our colleague and friend Steve Marshall, who collected the type series and took the photo of a living specimen (Fig. 1C View Figure 1 ), and from the Latin noun ' icona ', meaning image, in honour of Steve’s remarkable skills in natural history photography. The name should be treated as a feminine noun.