Omphreoides ranomafanae Kavanaugh and Rainio, 2016

Kavanaugh, David H. & Rainio, Johanna, 2016, Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar, Proceedings of the California Academy of Sciences 63 (7), pp. 201-268 : 224

Omphreoides ranomafanae Kavanaugh and Rainio , sp. nov.

Figure 13

TYPE MATERIAL.— Holotype ( Figs.13A–B ), a female, in NMNH, labeled: “ MADAGASCAR: Prov. Fianarantsoa, 7 km W Ranomafana , 900 m 17-22 February 1990 W. E. Steiner ”/ “at black light in montane rainforest near river and stream”/ “ HOLOTYPE Omphreoides ranomafanae Kavanaugh & Rainio sp. n. 2015” [red label].

TYPE LOCALITY.— Madagascar, Fianarantsoa Province, Ranomafana National Park .

DERIVATION OF SPECIES NAME.— The species epithet, ranomafanae , is a noun in apposition, derived from the name of the national park in which the type was collected.

RECOGNITION.— Size larger than average for genus, SBL = 13.4 mm. The unique holotype female of O. ranomafanae ( Fig. 13A ) is clearly unlike members of any described species of Omphreoides , with several features intermediate between the four typical species and O. quodi . The head of the holotype ( Fig. 13C ) is longer and more slender than that of females of any other species except O. quodi , which has an even longer and narrower head (see Jeannel 1949, Fig. 364). In O. ranomafanae , the tempora are straight, parallel in anterior their two-thirds, slightly more than twice as long as the diameter of eye, distinctly delimited posteriorly by narrowly round angles. The dorsal longitudinal grooves typical of all members of this genus, except those of Omphreoides bispinus Fairmaire (1896) , are extended as sharply-defined grooves only about halfway from the point of insertion of the anterior supraorbital setae to that of posterior supraorbital setae and not continued on to base of the head as they are in Omphreoides bucculentus Alluaud (1899a) and Omphreoides distinctus Alluaud (1936) . Posterior to the sharply-defined segments, broad and shallow depressions extend posteriorly, arcuately convergent and nearly joined in the midline near the back of the head. The pronotum ( Fig. 13C ) is subquadrate, longer than wide (ratio PW/PL = 0.75), with a wavy diagonal row of sparse, coarse punctures (seen also in O. distinctus and O. quodi members but absent from those of O. bucculentus and O. bispinus ) extended from near the anterior transverse impression paramedially to the basal foveae posteriorly; lateral pronotal margins with short but distinct sinuations anterior to slightly obtuse hind angles; median longitudinal impression deep, wider than in all other Omphreoides except O. quodi (in which the impression is even wider) and sparsely but coarsely punctate; lateral explanation narrow (narrower at middle than that in O. bucculentus members and similar to that in O. distinctus member. The elytra are relatively wider than in all other species; each elytron with a straight apical spine in line with interval 3.

GEOGRAPHICAL DISTRIBUTION.— At present, known only from the type locality.

HABITAT DISTRIBUTION.— The unique holotype was collected at ultraviolet light in montane rainforest near the junction of the Namorona River and a small tributary stream.

Figures

FIGURE 13. Digital images of holotype female of Omphreoides ranomafanae Kavanaugh & Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C. Forebody, dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.25 mm