Kamabrachys, Constant, 2023

Constant, Jérôme, 2023, Revision of the Eurybrachidae (XVII). The new Australian genus Kamabrachys gen. nov. with ten new species (Hemiptera: Fulgoromorpha), European Journal of Taxonomy 895 (1), pp. 1-133 : 109-116

publication ID

https://doi.org/ 10.5852/ejt.2023.895.2289

publication LSID

lsid:zoobank.org:pub:75CAAC73-8100-4D16-B970-4A533DBC7000

DOI

https://doi.org/10.5281/zenodo.8407990

persistent identifier

https://treatment.plazi.org/id/0A6687A0-9948-4F15-EA7C-FB6BFE4CFADD

treatment provided by

Plazi

scientific name

Kamabrachys
status

 

Kamabrachys v-carinatum gen. et sp. nov.

urn:lsid:zoobank.org:act:E3048362-A574-4420-96DF-6D92D901C636

Figs 5B View Fig , 69–72 View Fig View Fig View Fig View Fig

Platybrachys signata View in CoL – Lallemand 1935: 675 [recorded from Burnside, Northern Territory; misidentification!].

Diagnosis

The species can be recognized by the following combination of characters:

(1) posterior margin of the lateroventral lobes of the pygofer (lvl) rounded in lateral view ( Fig. 70A View Fig ) and rounded in ventral view ( Fig. 70C View Fig ), inner margin of lobes concave in ventral view ( Fig. 70C View Fig ), ventral margin rounded in caudal view ( Fig. 70E View Fig );

(2) posterior margin of laterodorsal process of gonostylus (ldg) roundly emarginate, leaving a rather widely open angle between laterodorsal process and ventral process of gonostylus (vp) in lateral view ( Fig. 70A View Fig );

(3) crest of furcate process of periandrium (cfp) along distal portion, merging basally and continued on basal portion ( Fig. 72D View Fig ), in lateral view, shortly tapering in distal portion ( Fig. 72E View Fig );

(4) laterodorsal process of periandrium (ldp) abruptly tapering after basal half and gently tapering in distal portion, distal half sinuate and curved laterodorsad, shorter than lateroventral process (lvp) ( Fig. 72D–E View Fig );

(5) lateroventral process of periandrium (lvp) dorsoventrally flattened, falcate in a long basal portion then twisted dorsolaterad distally with apex pointing dorsolaterocephalad ( Fig. 72D–F View Fig );

(6) apicodorsal process of aedeagus (adp) rather weakly sinuate, moderately wide distally and tapering towards the posterior, shorter than apicoventral process (avp) and quite strongly diverging from it

( Fig. 72A–C View Fig ); (7) posterior wing with weakly marked whitish marking along sutural margin ( Fig. 69A View Fig ).

Etymology

The species epithet is formed from the letter ‘v’ and the Latin adjective ‘ carinatus ’, meaning ‘crested, carinate’. It refers to the crests of the arms of the median furcate process of the periandrium which merge together on the shaft of the process in the shape of a ‘v’. The name is a noun phrase in which the adjectival portion of the name must be neuter, as all letters are grammatically neuter.

Type material

Holotype AUSTRALIA • ♂; [Northern Territory], Brock’s Creek ; [13°28′00″ S, 131°25′07″ E]; 8 May 1932; T. Campbell leg.; “Brock’s Ck, N.T., 8.5.32, T. Campbell”; ANIC. GoogleMaps

Paratypes AUSTRALIA • 1 ♂; [Northern Territory], 128 km S of Darwin; [13°30′19″ S, 131°22′50″ E]; 11 Jan. 1964; J. Sedlacek leg.; “Australia: N.T., 128 km S Darwin, 11.I.1964 ” “J. Sedlacek Collector, Bishop Museum”; BPBM GoogleMaps 1 ♂; [Northern Territory], Burnside ; [13°28′45″ S, 131°25′00″ E]; Apr. 1931; Handschin leg.; “Burnside, N.T., April 1931. Handschin” “ Platybrachys signata Dist. variety, det. W.E. China. 1934.”; FSAG GoogleMaps 1 ♂; [Northern Territory], Adelaide River Station ; [13°14′37″ S, 131°06′46″ E]; Apr. 1931; Handschin leg.; “Adelaide R. Stn, N.T., April 1931. Handschin”; NHMB GoogleMaps 1 ♂; [Northern Territory], Darwin ; [12°26′17″ S, 130°50′28″ E]; Jun. 1931; Handschin leg.; “Darwin, N.T., Juni 1931, Handschin”; NHMB GoogleMaps 1 ♂; [Northern Territory], Burnside ; [13°28′45″ S, 131°25′00″ E]; May 1932; Handschin leg.; “Burnside, N.T., April 1931. Handschin” “ Platybrachys signata Dist. var.”; NHMB GoogleMaps .

Description

MEASUREMENTS AND RATIOS. LT: ♂ (n = 6): 9.0 mm (8.64–9.24); BV/LV (n = 1): 3.35; BF/LF (n = 1): 1.25; LP+LM/BT (n = 1): 0.90; LM/LP (n = 1): 3.10; LTg/BTg (n = 1): 2.52; LW/BW (n = 1): 1.68.

HEAD ( Fig. 69A–D View Fig ). Slightly narrower than thorax. Vertex brown irregularly, usually rather poorly variegated with black; about 3.4 times as broad as long, concave with all margins slightly carinate; anterior and posterior margins rounded, parallel. Frons brown, sometimes with weak reddish hue, densely variegated with black, strongly more black than vertex; usually slightly paler on ventral portion, sometimes with a short paler median line on middle of disc; about 1.25 times as broad as long, slightly convex, slightly rugulose with peridiscal carina slightly marked; upper margin of frons weakly rounded, nearly straight in perpendicular view of frons; lateral margin slightly incurved above lateral projection of frons, more or less straight, oblique, under the projection. Clypeus brown, sometimes with weak reddish hue, with dark brown to black median line and regular oblique lines on each side of postclypeus, anteclypeus dark brown, black on sides, with median pale brown, sometimes slightly reddish line on carina; clypeus reaching mesocoxae, elongate, with median carina on anteclypeus prolongated with obsolete carina on postclypeus; base of clypeus rounded. Labium dark brown, sometimes slightly reddish, reaching metacoxae, with apical segment as long as broad, acuminate, shorter and slightly more slender than penultimate. Genae pale yellowish brown. Ocelli absent. Antennae brown, not surpassing eye, not visible from above; scape about as long as broad, pedicel subcylindrical, elongate, narrowing towards apex.

THORAX ( Fig. 69A, C View Fig ). About 1.10 times as broad as combined length of pro- and mesonotum. Pronotum brown irregularly variegated with black, generally as pale as vertex; disc weakly wrinkled, carina parallel to anterior margin and 2 obsolete impressed points on disc marked by black spot; paranotal lobes brown. Mesonotum brown irregularly variegated with black, more densely black and generally darker than pronotum; disc weakly wrinkled, median and peridiscal carinae weakly marked; median carina stopped before scutellum. Tegulae bicolour with ventral half mostly black and dorsal half pale brown.

TEGMINA ( Fig. 69A–C View Fig ). Pale brown irregularly variegated with darker brown and with black-brown markings along costal margin, slightly paler after nodal line than dark brown in distal portion; anteapical transverse white line usually interrupted in middle portion, when complete, very narrow in middle; small apical white marking along apical margin, extending on 2–4 cells; subrectangular black-brown transverse marking in middle portion of clavus extending from sutural margin to claval fold. Flat, elongate, about 2.5 times as long as broad; costal margin weakly rounded along basal fifth, then straight to nodal line; apex rounded.

VENATION. Veins ScP+RA and RP separated close to base; first fork of MP very basal, at the level of ScP+RA–RP separation; first fork of CuA slightly before apex of clavus; clavus closed; Pcu and A1 fused at about ¾ of clavus length; Pcu+A1 reaching apical angle of clavus; numerous cells along posterior half of costal margin; numerous cross-veins on apical third delimitating small elongate cells.

POSTERIOR WINGS ( Fig. 69A–B View Fig ). Yellow with apical ⅓ blackish brown, moderately broad band along posterior margin and anal area brown, paler than apical area; apical white marking sometimes present, weakly marked and extending on 2–3 cells; whitish marking along sutural margin between A1 and cubital fold, sometimes rather weakly marked. Colour more or less contrasted depending on individuals. Wings well developed, as broad as tegmina, about 1.7 times as long as broad, somewhat roundly truncate apically; anal area well developed; sutural margin hardly trilobous; not reaching apex of tegmina at rest. All main veins visible from base, forked at or after nodal line and forming a few closed cells; few transverse veinlets delimiting elongate cells on apical ¼; veins A1 and A2 sometimes with 2 terminals.

LEGS ( Fig. 69A–D View Fig ). Pro- and mesofemora and tibiae brown densely variegated with dark brown to black, dorsum of pro- and mesotibiae uniformly brown; pro- and mesotarsi paler than corresponding tibia but blackish apically; posterior legs brown with apex of tibial and tarsal spines darker. Pro- and mesofemora and tibiae dorsoventrally flattened, elongate and slender; metatibiae with 3 lateral and 10 apical spines; first metatarsomere ventrally with darker pad of microsetae at internoapical angle and two rows of 5 spines. Metatibiotarsal formula: (3) 10/4/0.

MALE TERMINALIA ( Figs 70–72 View Fig View Fig View Fig ). Pygofer (Py) rather short, as high as long and oblique in lateral view ( Fig. 70A View Fig ), about 2.6 times as wide as high in caudal view ( Fig. 70E View Fig ); basal apodeme (bap) well developed and sclerotized, sinuate in dorsal view ( Fig. 70A–C View Fig ); posterior margin abruptly emarginate on dorsal 1 / 5 ( Fig. 70A View Fig ), dorsal portion narrow ( Fig. 70A–B View Fig ); lateroventral lobe (lvl) rather strongly developed, projecting posterolaterally and broadly rounded in dorsal view ( Fig. 70B View Fig ); in ventral view, rounded anteriorly and posteriorly, more broadly rounded laterally and with inner margin excavate ( Fig. 70C View Fig ); in lateral strongly, roundly inflated posterad with uppermost portion sinuate, ventral margin more or less straight, horizontal, and anterior margin rounded ( Fig. 70A View Fig ); ventral portion separated from lateroventral lobe by a well-marked depression ( Fig. 70C, E View Fig ); ventral lamina (vlp) about 3.4 times as broad as long with anterior and posterior margins subparallel in ventral view, posterior margin weakly bisinuate, and posterolateral angles obliquely rounded laterally and weakly projecting posterad ( Fig. 70C View Fig ). Anal tube 1.1 times as broad as long (An), dorsoventrally flattened ( Fig. 70A View Fig ), subcircular with lateral margins more or less regularly rounded; apical margin rather strongly indented in middle ( Fig. 70B View Fig ); paraprocts (pa) at basal ¼, well developed. Gonostyli (G) rather short, 1.47 times as broad as long in ventral view ( Fig. 70C View Fig ); ventral process (vp) in ventral view rather short, subtriangular with lateral margin rather strongly sinuate, then rounded apically ( Fig. 70C View Fig ); laterodorsal process (ldg) narrowly visible on sides in ventral view ( Fig. 70C View Fig ), strongly sinuate in caudal view with mediolateral angle (mla) strongly marked ( Fig. 70E View Fig ), mediolateral angle projecting laterally ( Fig. 70A, D View Fig ), in lateral view, posterior margin roundly concave under mediolateral angle ( Fig. 70A View Fig ); dorsal hook (dhg) more or less subtriangular in lateral view, with anterior margin roundly excavate and posterior margin rounded above basal indentation ( Fig. 70A View Fig ), and apically curved laterodorsad and slightly cephalad ( Fig. 70B, E View Fig ). Connective (cv) strongly developed, about 1.3 times as long as aedeagus s. lat., with anterior pair of lateral apodemes forming a moderately broad lateral lamina directed posterolaterad and rounded apically; posterior pair of apodemes ribbon-like, strongly developed laterally and sinuate ( Fig. 71A–D View Fig ). Aedeagus s. lat. moderately upcurved with periandrium reaching about same level as aedeagus s. str. posterad ( Fig. 71E–I View Fig ). Aedeagus s. str. ( Fig. 72A–C View Fig ) with apical processes about 7.0 times as long as broad at mid-length; apicoventral process (avp) smoothly tapering towards the posterior and narrowly rounded apically ( Fig. 72C View Fig ); apicodorsal process (adp) in dorsal view ( Fig. 72A View Fig ) moderately sinuate and moderately inflated, rather regularly tapering towards apex, distally slightly diverging from apicoventral process and slightly shorter than the latter. Dorsal periandrium ( Fig. 72D–F View Fig ) with laterodorsal processes (ldp) dorsoventrally laminate basally and weakly curved laterad, then abruptly tapering, reflexed and curved laterodorsad along distal half, sinuate in dorsal view and apically pointed; lateroventral processes (lvp) dorsoventrally flattened, directed laterad, oboval on basal + / 5, broadening from base to first ⅓, then gently tapering up to abruptly narrowing distal 1 / 5, distal 1 / 5 directed laterocephalad and first slightly curved dorsad, then ventrad towards pointed apex, projecting posterolaterally beyond laterodorsal processes; basomedian processes (bmp) apically roundly blunt in dorsal view and with sides parallel in ventral view; median furcate process (mfp) with furca slightly shorter than shaft ( Fig. 71I View Fig ) and rather strongly upcurved in lateral view ( Fig. 72E View Fig ), dorsal crest (cfp) of arms of furca strongly developed, merging into a single crest extending to half-length of shaft, crests on arms slightly reflexed laterad in dorsal view ( Fig. 72D View Fig ), crest rather high along most of its length and with dorsal margin irregularly crenelated, tapering distally in lateral view ( Fig. 72E View Fig ), inner lateral walls of furca granulose ( Fig. 72D View Fig ).

SEXUAL DIMORPHISM. Unknown (no female specimen available).

Distribution and biology

The species is currently recorded from five nearby locations in northeastern Northern Territory ( Fig. 5B View Fig ). Bioregions: Darwin Coastal and Pine Creek.

The specimens of the type series were collected during the months of January, and April to June.

Host plant unknown.

FSAG

Belgium, Gembloux, Zoologie Generale et Appliquee

NHMB

Switzerland, Basel, Naturhistorisches Museum

ANIC

Australian National Insect Collection

BPBM

Bishop Museum

FSAG

Faculte des Sciences Agronomiques de Gembloux

NHMB

Natural History Museum Bucharest

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Hemiptera

Family

Eurybrachidae

Loc

Kamabrachys

Constant, Jérôme 2023
2023
Loc

Platybrachys signata

Lallemand V. 1935: 675
1935
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