Theodoxus gloeri Odabasi & Arslan, 2015

Sands, Arthur F, Gloeer, Peter, Guerlek, Mustafa E, Albrecht, Christian & Neubauer, Thomas A, 2020, A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species, Zoosystematics and Evolution 96 (1), pp. 25-66 : 25

Theodoxus gloeri Odabasi & Arslan, 2015 Figure 11A-C

Theodoxus gloeri Odabaşı and Arslan 2015: 329-330, fig. 2; Gürlek et al. 2019: 2993; Glöer 2019: 44, fig. 29.

Type locality.

Balıkdamı Wetland-Sakarya River, Eskişehir, Turkey; 39.15277°N, 31.61562°E (Fig. 3D ).

Type material.

Holotype and seven paratypes are deposited in COMULM (coll. no. COMULM-G 0052-0053) . Further nine paratypes are deposited in OGUHB (coll. no. OGUHB-01759) . An additional paratype is stored in the private collection of one of us (coll. Glöer; Fig. 11A-C ) .

Remarks.

Theodoxus gloeri Odabaşı & Arslan, 2015 shows strong similarity to the fossil species T. pilidei ( Tournouër, 1879) from Romania and T. lamelliferus (Milaschewitsch, 1912) from the Black Sea (near Alushta, Crimea) with strongly ribbed shells, but it is smaller than both ( Odabaşı and Arslan 2015; Glöer 2019; Fig. 11 ). Although it is expected to be an extant subterranean species, no living material has ever been recorded and its phylogenetic position is presently unknown ( Odabaşı and Arslan 2015).

Distribution.

Only known from two sites in very close proximity within the Balıkdamı Wetland-Sakarya River, Eskişehir, Turkey ( Odabaşı and Arslan 2015; Fig. 3D ).

Figures

Figure 3. Occurrence map for Theodoxus spp. in Asia. All points conform to specimens used in the phylogenetic analyses (Suppl. material 1: Table S 1), except for T. gloeri for which no molecular data are available and only the type and paratype localities are indicated. Species are partitioned across four identical maps (A - D) to limit overlap of points as far as possible. Barring T. gloeri, closer related species are grouped together (Fig. 2).

Figure 11. Theodoxus gloeri Odabasi & Arslan, 2015. A - C. Paratype from Balikdami Wetland-Sakarya River, Eskisehir, Turkey (coll. Gloeer). Scale bar: 1 mm.