Pseudochillus (Kaszabochillus) helferi 2015

Fouquè, René, 2015, A review of the genera Indostola, Tagenostola, Indochillus, Pseudethas, and Pseudochillus gen. nov. in South East Asia (Coleoptera: Tenebrionidae: Stenosini), Acta Entomologica Musei Nationalis Pragae 55 (1), pp. 217-242 : 229-230

Pseudochillus (Kaszabochillus) helferi sp. nov.

( Figs 9 , 14 , 21 , 33 )

Type locality. Myanmar, Tenasserim.

Type material. HOLOTYPE: ♀, ‘TENASSERIM / Coll. Helferʼ ( NMPC).

Description. Body length 3.9 mm, body width 1.1 mm. Body, legs and antennae rusty brown.

Head length/width ratio 1.06, head widest at anterior margin of eyes. Tempora widest at posterior margin of eyes, from there slowly narrowed up to distance equal to half eye length, than rounded and sharply narrowed to cervix. Cervical constriction on the same level as interior margin of eyes. Genae arched from posterior margin of eyes to the widest place, there arched and directly narrowed to clypeus. Clypeus straight without tooth. Inner margin of eye with reduced suborbital keel vanishing in middle of eye groove. Cervical constriction concave. Frons with two distinct impressions. Occiput to cervical constriction very slightly flattened. Ratio of head/cervix widths 1.73. Eyes ( Fig. 33 ) completely divided by genae, dorsal part with 18 facets in 3 rows, ventral part with 7 facets arranged in triangle. Punctation teardrop like, distance between punctures as large as their diameter, on occiput punctures rounded and largest, almost joined. Punctures gradually smaller towards clypeus, with very small setae as long as punctures diameter, oriented forwards. Clypeus with longer setae and longer impressions. Space between punctures finely wrinkled. Antennae ( Fig. 21 ) with yellow setae directing forwards. Setae cut apically and there widest, on middle antennomeres as long as 0.4 length of these antennomeres. Antennomeres 2–11 combined twice as long as pronotum width; antennomeres trapezoidal, last antennomere cut latero-apically; antennomere 3 longest, 1.3 times longer than antennomere 4, antennomeres 9 and 10 widest; length ratio of antennomeres 2–11 100: 135: 104: 96: 96: 96: 96: 104: 104: 104, width ratio 109: 126: 126: 117: 122: 122: 126: 139: 139: 130.

Pronotum longer than wide (1.22), widest in anterior fifth; width ratio of head / anterior edges of pronotum / widest point of pronotum / posterior edges of pronotum 100: 75: 87: 62. Anterior corners obtusely rounded, not protruding. Posterior corners obtuse. Pronotal base convex. Anterior margin straight. Lateral margin cordiform. Pronotum with mid longitudinal impression in its whole length. Pronotum in rear view arched with impression in middle and with impression on each side (the latter is therefore concave). Punctation dense and larger than that on head, rounded. Punctures largest near lateral margins. Space without punctures at distance equal to 1.5–2.0 diameter of lateral punctures from lateral margin. Punctures with forward-oriented yellow setae at least as long as puncture diameter. Space between punctures very finely wrinkled.

Elytra elongate; 2.8 times longer and 1.6 times wider than pronotum, elytra length/width ratio 2.20. Base of elytra concave, narrower than pronotal base (0.84). Each elytron with 10 rows of punctures, 8 on dorsal side, 2 on deflexed part; intervals 3, 5, 7 and 9 keeled. Intervals 5 and 9 flattened on base and elytral declivity. Intervals 3 and 7 joined before apex. Junction of interval 7 and 9 forming small obliquely directed humeral corner ( Fig. 14 ). Punctures deep, rounded and larger than those on pronotum. Distance between punctures in row smaller than puncture diameters. Interspaces gently wrinkled. Elytra without setae. Epipleura with one row of punctures throughout its whole length, its punctures somewhat smaller than in row 10, becoming smaller towards apex. Humeral calli not developed, apterous.

Legs finely covered with pale adherent hairs. Female tibiae without small inner subapical tooth (male tibiae unknown). Width/length ratio of posterior tibia 0.13.

Abdomen with small, rounded punctures gradually smaller towards apex. Distance between punctures on first ventrite 3–4 times higher than puncture diameters; on second ventrite 2 times higher than puncture diameters; on third and fourth ventrites 1.0–1.5 times higher than puncture diameters; and on anal ventrite with space between punctures equal to puncture diameter.

Male unknown.

Differential diagnosis. This species is very similar to P. (K.) andamanus ( Kaszab, 1981) but at first sight it differs in presence of mid-longitudinal impression on pronotum and in impunctate lateral margins of pronotum.

Etymology. The new species is named in honour of Johann Wilhelm Helfer (1810–1840), Czech doctor, traveller, and entomologist (see BEZDĚK & HÁJEK 2010), collector of the type species.

Distribution. Myanmar: Tenasserim.

Remarks. According to BEZDĚK & HÁJEK (2010) the holotype was collected between the years 1837–1840.

Figures

Figs 5–9. Dorsal view. 5 – Indochillus cristatus Koch, 1941, holotype, India (Karnakata). 6 – Pseudochillus (Pseudochillus) bangaloreanus (Kaszab, 1981), holotype ♂, India (Bangalore). 7 – P. (P.) aalbui sp. nov., holotype ♂, India (Tamil Nadu).8 – P.(Kaszabochillus) andamanus (Kaszab, 1981), paratype♀, India (Andaman Islands). 9 – P. (K.) helferi sp. nov, holotype ♀, Myanmar (Tennasserim). Scale bar = 4 mm.

Figs 10–16. 10–12 – dorsal view:10 – Pseudochillus (Micropseudochillus) palawanus sp.nov., holotype♂, Philippines (Palawan); 11 – P.(M.) thailandicus sp. nov., holotype ♂, Thailand.; 12 – P.(M.) indochinensis sp. nov., paratype ♀, Vietnam. 13 – P.(M.) indochinensis sp. nov., tooth on clypeus; 14–15 – humeral corner: 14 – P.(Kaszabochillus) helferi sp. nov.; 15 – P.(Pseudochillus) bangaloreanus (Kaszab, 1981). 16 – P.(P.) bangaloreanus (Kaszab, 1981), punctures on epipleura. Scale bar = 4 mm (Figs 10–12).

Figs 17–26. Antenna. 17 – Indochillus cristatus Koch, 1941. 18 – Pseudochillus (Pseudochillus) bangaloreanus (Kaszab, 1981). 19 – P.(P.) aalbui sp. nov. 20 – P.(Kaszabochillus) andamanus (Kaszab, 1981). 21 – P.(K.) helferi sp.nov. 22 – Indostola kaengkrachanensis sp.nov. 23 – I.pulchella Medvedev, 1991.24 – Pseudochillus (Micropseudochillus) palawanus sp. nov. 25 – P.(M.) thailandicus sp. nov. 26 – P.(M.) indochinensis sp. nov. Scale bar = 1 mm.

Figs 27–36. Lateral view of head with eyes. 27 – Indostola pulchella Medvedev, 1991. 28 – I. kaengkrachanensis sp. nov.29 – Indochillus cristatus Koch, 1941. 30 – Pseudochillus (Pseudochillus) bangaloreanus (Kaszab, 1981). 31 – P. (P.) aalbui sp. nov. 32 – P.(Kaszabochillus) andamanus (Kaszab, 1981). 33 – P.(K.) helferi sp. nov. 34 – Pseudochillus (Micropseudochillus) palawanus sp. nov. 35 – P.(M.) thailandicus sp. nov. 36 – P.(M.) indochinensis sp. nov.