Halys, Fabricius, 1803

Salini, S., 2019, Revision of the genus Halys (Hemiptera: Pentatomidae) with description of a new species from India, Zootaxa 4586 (2), pp. 351-375 : 370

Key to species of females of Halys from India

1. Valvifers IX with posterior margin incised medially giving a medially divided appearance ( Figs 56 , 88 ); valvulae VIII with apical (= caudal) margin uniformly rounded ( Figs 54, 55 , 90 ).................................................. 2

1'. Valvifers IX with posterior margin not incised medially, either uniformly rounded or concave ( Figs 40 , 74, 75 );valvulae VIII with apical (= caudal) margin not uniformly rounded rather slightly concave medially ( Figs 41 , 76 ).................... 3

2. Posterior margin of valvifers VIII undulate and with distinct concavity adjacent to inner posterior angles and mesal margin straight ( Figs 55, 56 )................................................................. H. serrigera Westwood

2' Posterior margin of valvifers VIII convex to straight and mesal margin slightly concave ( Figs 88, 89 ).. H. sulcata (Thunberg)

3. Pronotum broadly angulate, disc slightly elevated, scutellum black ( Figs 1, 3 , 15, 17 )............. H. mudigerensis sp. nov.

3'. Pronotum narrowly angulate, disc more or less flat, scutellum concolourous withdorsal body ( Fig. 6 )...... H. shaista Ghauri

Figures

FIGURES 49–56. H. serrigera Westwood, 1837—male and female genitalia. 49, phallus (ventral); 50, phallus (dorsal); 51–52, phallus (lateral); 53, spermatheca; 54, valvulae VIII; 55, female terminalia (before digestion); 56, female terminalia (after digestion). Lettering: ad—aedeagus; cp—capitate processes; dcp—dorsal conjunctival process; pa—processes of aedeagus; lt VIII—laterotergite VIII; lt IX— laterotergite IX; sp—spermathecal pump; vlf VIII—valvifers VIII; vlf IX—valvifers IX.

FIGURES 84–90. H. sulcata (Thunberg, 1783)—male and female genitalia. 84, phallus (dorsal); 85, phallus (ventral); 86, phallus (lateral); 87, spermatheca; 88, female terminalia (before digestion); 89, female terminalia (after digestion); 90,valvulae VIII.

FIGURES 36–41. H. mudigerensis sp. nov.—male and female genitalia. 36, phallus (dorsal); 37, phallus (ventral); 38, phallus (lateral); 39, spermatheca; 40, female terminalia; 41,valvulae VIII. Lettering: dcp—dorsal conjunctival processes; lt VIIIlaterotergite VIII; lt IX—laterotergite IX; pa—processes of aedeagus; sp—spermathecal pump; vcp—ventral conjunctival process; vlf VIII—valvifers VIII; vlf IX—valvifers IX.

FIGURES 69–76. H. shaista Ghauri, 1988—male and female genitalia. 69, phallus (dorsal); 70, phallus (ventral); 71–72, phallus (lateral); 73, spermatheca; 74, female terminalia (before digestion); 75, female terminalia (after digestion); 76, valvulae VIII.

FIGURES 1–9. Halys Fabricius, 1803 (dorsal and ventral habitus). 1–2, Halys mudigerensis sp. nov. (male); 3, H. mudigerensis sp. nov. (female without abdomen); 4–5, H. serrigera Westwood, 1837; 6–7, H. shaista Ghauri, 1988; 8–9, H. sulcata (Thunberg, 1783)

FIGURES 10–17. Halys Fabricius, 1803.10, H. mudigerensis sp. nov. (head—dorsal);11, H. serrigera Westwood, 1837 (head—dorsal); 12, H. serrigera Westwood, 1837 (head—ventral); 13, H. shaista Ghauri, 1988 (head—dorsal); 14, H. sulcata (Thunberg, 1783) (head—dorsal); 15, H. mudigerensis sp. nov. (scutellum); 16, H. serrigera Westwood, 1837 (head—lateral); 17, H. mudigerensis sp. nov. (pronotum). Lettering: at—antenniferous tubercle; b—bucculae; lr—labrum; mt—tooth on lateral margin of mandibular plates.